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624_19 | Starting in 1982, dividends from the fund's annual growth have been paid out each year to eligible Alaskans, ranging from an initial $1,000 in 1982 (equal to three years' payout, as the distribution of payments was held up in a lawsuit over the distribution scheme) to $3,269 in 2008 (which included a one-time $1,200 "Resource Rebate"). Every year, the state legislature takes out 8% from the earnings, puts 3% back into the principal for inflation proofing, and the remaining 5% is distributed to all qualifying Alaskans. To qualify for the Permanent Fund Dividend, one must have lived in the state for a minimum of 12 months, maintain constant residency subject to allowable absences, and not be subject to court judgments or criminal convictions which fall under various disqualifying classifications or may subject the payment amount to civil garnishment.
The Permanent Fund is often considered to be one of the leading examples of a basic income policy in the world.
Cost of living
The cost of goods in Alaska has long been higher than in the contiguous 48 states. Federal government employees, particularly United States Postal Service (USPS) workers and active-duty military members, receive a Cost of Living Allowance usually set at 25% of base pay because, while the cost of living has gone down, it is still one of the highest in the country.
Rural Alaska suffers from extremely high prices for food and consumer goods compared to the rest of the country, due to the relatively limited transportation infrastructure.
Agriculture and fishing
Due to the northern climate and short growing season, relatively little farming occurs in Alaska. Most farms are in either the Matanuska Valley, about northeast of Anchorage, or on the Kenai Peninsula, about southwest of Anchorage. The short 100-day growing season limits the crops that can be grown, but the long sunny summer days make for productive growing seasons. The primary crops are potatoes, carrots, lettuce, and cabbage. | [
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624_20 | The Tanana Valley is another notable agricultural locus, especially the Delta Junction area, about southeast of Fairbanks, with a sizable concentration of farms growing agronomic crops; these farms mostly lie north and east of Fort Greely. This area was largely set aside and developed under a state program spearheaded by Hammond during his second term as governor. Delta-area crops consist predominantly of barley and hay. West of Fairbanks lies another concentration of small farms catering to restaurants, the hotel and tourist industry, and community-supported agriculture.
Alaskan agriculture has experienced a surge in growth of market gardeners, small farms and farmers' markets in recent years, with the highest percentage increase (46%) in the nation in growth in farmers' markets in 2011, compared to 17% nationwide. The peony industry has also taken off, as the growing season allows farmers to harvest during a gap in supply elsewhere in the world, thereby filling a niche in the flower market.
Alaska, with no counties, lacks county fairs. However, a small assortment of state and local fairs (with the Alaska State Fair in Palmer the largest), are held mostly in the late summer. The fairs are mostly located in communities with historic or current agricultural activity, and feature local farmers exhibiting produce in addition to more high-profile commercial activities such as carnival rides, concerts and food. "Alaska Grown" is used as an agricultural slogan.
Alaska has an abundance of seafood, with the primary fisheries in the Bering Sea and the North Pacific. Seafood is one of the few food items that is often cheaper within the state than outside it. Many Alaskans take advantage of salmon seasons to harvest portions of their household diet while fishing for subsistence, as well as sport. This includes fish taken by hook, net or wheel.
Hunting for subsistence, primarily caribou, moose, and Dall sheep is still common in the state, particularly in remote Bush communities. An example of a traditional native food is Akutaq, the Eskimo ice cream, which can consist of reindeer fat, seal oil, dried fish meat and local berries.
Alaska's reindeer herding is concentrated on Seward Peninsula, where wild caribou can be prevented from mingling and migrating with the domesticated reindeer. | [
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624_21 | Most food in Alaska is transported into the state from "Outside" (the other 49 US states), and shipping costs make food in the cities relatively expensive. In rural areas, subsistence hunting and gathering is an essential activity because imported food is prohibitively expensive. Although most small towns and villages in Alaska lie along the coastline, the cost of importing food to remote villages can be high, because of the terrain and difficult road conditions, which change dramatically, due to varying climate and precipitation changes. The cost of transport can reach as high as 50¢ per pound ($1.10/kg) or more in some remote areas, during the most difficult times, if these locations can be reached at all during such inclement weather and terrain conditions. The cost of delivering a of milk is about $3.50 in many villages where per capita income can be $20,000 or less. Fuel cost per gallon is routinely twenty to thirty cents higher than the contiguous United States average, with only Hawaii having higher prices.
Culture
Some of Alaska's popular annual events are the Iditarod Trail Sled Dog Race from Anchorage to Nome, World Ice Art Championships in Fairbanks, the Blueberry Festival and Alaska Hummingbird Festival in Ketchikan, the Sitka Whale Fest, and the Stikine River Garnet Fest in Wrangell. The Stikine River attracts the largest springtime concentration of American bald eagles in the world.
The Alaska Native Heritage Center celebrates the rich heritage of Alaska's 11 cultural groups. Their purpose is to encourage cross-cultural exchanges among all people and enhance self-esteem among Native people. The Alaska Native Arts Foundation promotes and markets Native art from all regions and cultures in the State, using the internet.
Music
Influences on music in Alaska include the traditional music of Alaska Natives as well as folk music brought by later immigrants from Russia and Europe. Prominent musicians from Alaska include singer Jewel, traditional Aleut flautist Mary Youngblood, folk singer-songwriter Libby Roderick, Christian music singer-songwriter Lincoln Brewster, metal/post hardcore band 36 Crazyfists and the groups Pamyua and Portugal. The Man. | [
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624_22 | There are many established music festivals in Alaska, including the Alaska Folk Festival, the Fairbanks Summer Arts Festival the Anchorage Folk Festival, the Athabascan Old-Time Fiddling Festival, the Sitka Jazz Festival, and the Sitka Summer Music Festival. The most prominent orchestra in Alaska is the Anchorage Symphony Orchestra, though the Fairbanks Symphony Orchestra and Juneau Symphony are also notable. The Anchorage Opera is currently the state's only professional opera company, though there are several volunteer and semi-professional organizations in the state as well.
The official state song of Alaska is "Alaska's Flag", which was adopted in 1955; it celebrates the flag of Alaska.
Alaska in film and on television
Alaska's first independent picture entirely made in Alaska was The Chechahcos, produced by Alaskan businessman Austin E. Lathrop and filmed in and around Anchorage. Released in 1924 by the Alaska Moving Picture Corporation, it was the only film the company made.
One of the most prominent movies filmed in Alaska is MGM's Eskimo/Mala The Magnificent, starring Alaska Native Ray Mala. In 1932, an expedition set out from MGM's studios in Hollywood to Alaska to film what was then billed as "The Biggest Picture Ever Made". Upon arriving in Alaska, they set up "Camp Hollywood" in Northwest Alaska, where they lived during the duration of the filming. Louis B. Mayer spared no expense in spite of the remote location, going so far as to hire the chef from the Hotel Roosevelt in Hollywood to prepare meals.
When Eskimo premiered at the Astor Theatre in New York City, the studio received the largest amount of feedback in its history. Eskimo was critically acclaimed and released worldwide; as a result, Mala became an international movie star. Eskimo won the first Oscar for Best Film Editing at the Academy Awards, and showcased and preserved aspects of Inupiat culture on film. | [
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624_23 | The 1983 Disney movie Never Cry Wolf was at least partially shot in Alaska. The 1991 film White Fang, based on Jack London's 1906 novel and starring Ethan Hawke, was filmed in and around Haines. Steven Seagal's 1994 On Deadly Ground, starring Michael Caine, was filmed in part at the Worthington Glacier near Valdez. The 1999 John Sayles film Limbo, starring David Strathairn, Mary Elizabeth Mastrantonio, and Kris Kristofferson, was filmed in Juneau.
The psychological thriller Insomnia, starring Al Pacino and Robin Williams, was shot in Canada, but was set in Alaska. The 2007 film directed by Sean Penn, Into The Wild, was partially filmed and set in Alaska. The film, which is based on the novel of the same name, follows the adventures of Christopher McCandless, who died in a remote abandoned bus along the Stampede Trail west of Healy in 1992.
Many films and television shows set in Alaska are not filmed there; for example, Northern Exposure, set in the fictional town of Cicely, Alaska, was filmed in Roslyn, Washington. The 2007 horror feature 30 Days of Night is set in Barrow, Alaska, but was filmed in New Zealand.
Many reality television shows are filmed in Alaska. In 2011, the Anchorage Daily News found ten set in the state.
Public health and public safety
The Alaska State Troopers are Alaska's statewide police force. They have a long and storied history, but were not an official organization until 1941. Before the force was officially organized, law enforcement in Alaska was handled by various federal agencies. Larger towns usually have their own local police and some villages rely on "Public Safety Officers" who have police training but do not carry firearms. In much of the state, the troopers serve as the only police force available. In addition to enforcing traffic and criminal law, wildlife Troopers enforce hunting and fishing regulations. Due to the varied terrain and wide scope of the Troopers' duties, they employ a wide variety of land, air, and water patrol vehicles.
Many rural communities in Alaska are considered "dry", having outlawed the importation of alcoholic beverages. Suicide rates for rural residents are higher than urban. | [
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624_24 | Domestic abuse and other violent crimes are also at high levels in the state; this is in part linked to alcohol abuse. Alaska has the highest rate of sexual assault in the nation, especially in rural areas. The average age of sexually assaulted victims is 16 years old. In four out of five cases, the suspects were relatives, friends or acquaintances.
Education
The Alaska Department of Education and Early Development administers many school districts in Alaska. In addition, the state operates a boarding school, Mt. Edgecumbe High School in Sitka, and provides partial funding for other boarding schools, including Nenana Student Living Center in Nenana and The Galena Interior Learning Academy in Galena.
There are more than a dozen colleges and universities in Alaska. Accredited universities in Alaska include the University of Alaska Anchorage, University of Alaska Fairbanks, University of Alaska Southeast, and Alaska Pacific University. Alaska is the only state that has no institutions that are part of NCAA Division I.
The Alaska Department of Labor and Workforce Development operates AVTEC, Alaska's Institute of Technology. Campuses in Seward and Anchorage offer one-week to 11-month training programs in areas as diverse as Information Technology, Welding, Nursing, and Mechanics.
Alaska has had a problem with a "brain drain". Many of its young people, including most of the highest academic achievers, leave the state after high school graduation and do not return. , Alaska did not have a law school or medical school. The University of Alaska has attempted to combat this by offering partial four-year scholarships to the top 10% of Alaska high school graduates, via the Alaska Scholars Program.
Beginning in 1998, schools in rural Alaska must have at least 10 students to retain funding from the state, and campuses not meeting the number close. This was due to the loss in oil revenues that previously propped up smaller rural schools. In 2015, there was a proposal to raise that minimum to 25, but legislators in the state largely did not agree.
Transportation
Roads | [
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624_25 | Transportation
Roads
Alaska has few road connections compared to the rest of the U.S. The state's road system, covering a relatively small area of the state, linking the central population centers and the Alaska Highway, the principal route out of the state through Canada. The state capital, Juneau, is not accessible by road, only a car ferry; this has spurred debate over decades about moving the capital to a city on the road system, or building a road connection from Haines. The western part of Alaska has no road system connecting the communities with the rest of Alaska.
The Interstate Highways in Alaska consists of a total of . One unique feature of the Alaska Highway system is the Anton Anderson Memorial Tunnel, an active Alaska Railroad tunnel recently upgraded to provide a paved roadway link with the isolated community of Whittier on Prince William Sound to the Seward Highway about southeast of Anchorage at Portage. At , the tunnel was the longest road tunnel in North America until 2007. The tunnel is the longest combination road and rail tunnel in North America.
Rail
Built around 1915, the Alaska Railroad (ARR) played a key role in the development of Alaska through the 20th century. It links north Pacific shipping through providing critical infrastructure with tracks that run from Seward to Interior Alaska by way of South Central Alaska, passing through Anchorage, Eklutna, Wasilla, Talkeetna, Denali, and Fairbanks, with spurs to Whittier, Palmer and North Pole. The cities, towns, villages, and region served by ARR tracks are known statewide as "The Railbelt". In recent years, the ever-improving paved highway system began to eclipse the railroad's importance in Alaska's economy.
The railroad played a vital role in Alaska's development, moving freight into Alaska while transporting natural resources southward, such as coal from the Usibelli coal mine near Healy to Seward and gravel from the Matanuska Valley to Anchorage. It is well known for its summertime tour passenger service. | [
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624_26 | The Alaska Railroad was one of the last railroads in North America to use cabooses in regular service and still uses them on some gravel trains. It continues to offer one of the last flag stop routes in the country. A stretch of about of track along an area north of Talkeetna remains inaccessible by road; the railroad provides the only transportation to rural homes and cabins in the area. Until construction of the Parks Highway in the 1970s, the railroad provided the only land access to most of the region along its entire route.
In northern Southeast Alaska, the White Pass and Yukon Route also partly runs through the state from Skagway northwards into Canada (British Columbia and Yukon Territory), crossing the border at White Pass Summit. This line is now mainly used by tourists, often arriving by cruise liner at Skagway. It was featured in the 1983 BBC television series Great Little Railways.
The Alaska Rail network is not connected to Outside. (The nearest link to the North American railway network is the northwest terminus of the Canadian National Railway at Prince Rupert, British Columbia, several hundred miles to the southeast.) In 2000, the U.S. Congress authorized $6 million to study the feasibility of a rail link between Alaska, Canada, and the lower 48.
Some private companies provides car float service between Whittier and Seattle.
Marine transport
Many cities, towns and villages in the state do not have road or highway access; the only modes of access involve travel by air, river, or the sea.
Alaska's well-developed state-owned ferry system (known as the Alaska Marine Highway) serves the cities of southeast, the Gulf Coast and the Alaska Peninsula. The ferries transport vehicles as well as passengers. The system also operates a ferry service from Bellingham, Washington and Prince Rupert, British Columbia, in Canada through the Inside Passage to Skagway. The Inter-Island Ferry Authority also serves as an important marine link for many communities in the Prince of Wales Island region of Southeast and works in concert with the Alaska Marine Highway.
In recent years, cruise lines have created a summertime tourism market, mainly connecting the Pacific Northwest to Southeast Alaska and, to a lesser degree, towns along Alaska's gulf coast. The population of Ketchikan for example fluctuates dramatically on many days—up to four large cruise ships can dock there at the same time. | [
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624_27 | Air transport
Cities not served by road, sea, or river can be reached only by air, foot, dogsled, or snowmachine, accounting for Alaska's extremely well developed bush air services—an Alaskan novelty. Anchorage and, to a lesser extent Fairbanks, is served by many major airlines. Because of limited highway access, air travel remains the most efficient form of transportation in and out of the state. Anchorage recently completed extensive remodeling and construction at Ted Stevens Anchorage International Airport to help accommodate the upsurge in tourism (in 2012–2013, Alaska received almost two million visitors).
Regular flights to most villages and towns within the state that are commercially viable are challenging to provide, so they are heavily subsidized by the federal government through the Essential Air Service program. Alaska Airlines is the only major airline offering in-state travel with jet service (sometimes in combination cargo and passenger Boeing 737-400s) from Anchorage and Fairbanks to regional hubs like Bethel, Nome, Kotzebue, Dillingham, Kodiak, and other larger communities as well as to major Southeast and Alaska Peninsula communities.
The bulk of remaining commercial flight offerings come from small regional commuter airlines such as Ravn Alaska, PenAir, and Frontier Flying Service. The smallest towns and villages must rely on scheduled or chartered bush flying services using general aviation aircraft such as the Cessna Caravan, the most popular aircraft in use in the state. Much of this service can be attributed to the Alaska bypass mail program which subsidizes bulk mail delivery to Alaskan rural communities. The program requires 70% of that subsidy to go to carriers who offer passenger service to the communities.
Many communities have small air taxi services. These operations originated from the demand for customized transport to remote areas. Perhaps the most quintessentially Alaskan plane is the bush seaplane. The world's busiest seaplane base is Lake Hood, located next to Ted Stevens Anchorage International Airport, where flights bound for remote villages without an airstrip carry passengers, cargo, and many items from stores and warehouse clubs. | [
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624_28 | In 2006, Alaska had the highest number of pilots per capita of any U.S. state. In Alaska there are 8,795 active pilot certificates as of 2020.
Of these, there are 2,507 Private, 1,496 Commercial, 2,180 Airline Transport, and 2,239 Student. There are also 3,987 pilots with a Instrument rating and 1,511 Flight Instructors.
Other transport
Another Alaskan transportation method is the dogsled. In modern times (that is, any time after the mid-late 1920s), dog mushing is more of a sport than a true means of transportation. Various races are held around the state, but the best known is the Iditarod Trail Sled Dog Race, a trail from Anchorage to Nome (although the distance varies from year to year, the official distance is set at ). The race commemorates the famous 1925 serum run to Nome in which mushers and dogs like Togo and Balto took much-needed medicine to the diphtheria-stricken community of Nome when all other means of transportation had failed. Mushers from all over the world come to Anchorage each March to compete for cash, prizes, and prestige. The "Serum Run" is another sled dog race that more accurately follows the route of the famous 1925 relay, leaving from the community of Nenana (southwest of Fairbanks) to Nome.
In areas not served by road or rail, primary transportation in summer is by all-terrain vehicle and in winter by snowmobile or "snow machine", as it is commonly referred to in Alaska.
Data transport
Alaska's internet and other data transport systems are provided largely through the two major telecommunications companies: GCI and Alaska Communications. GCI owns and operates what it calls the Alaska United Fiber Optic system and, as of late 2011, Alaska Communications advertised that it has "two fiber optic paths to the lower 48 and two more across Alaska. In January 2011, it was reported that a $1 billion project to connect Asia and rural Alaska was being planned, aided in part by $350 million in stimulus from the federal government.
Law and government
State government | [
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] |
624_29 | Law and government
State government
Like all other U.S. states, Alaska is governed as a republic, with three branches of government: an executive branch consisting of the governor of Alaska and his or her appointees which head executive departments; a legislative branch consisting of the Alaska House of Representatives and Alaska Senate; and a judicial branch consisting of the Alaska Supreme Court and lower courts.
The state of Alaska employs approximately 16,000 people statewide.
The Alaska Legislature consists of a 40-member House of Representatives and a 20-member Senate. Senators serve four-year terms and House members two. The governor of Alaska serves four-year terms. The lieutenant governor runs separately from the governor in the primaries, but during the general election, the nominee for governor and nominee for lieutenant governor run together on the same ticket.
Alaska's court system has four levels: the Alaska Supreme Court, the Alaska Court of Appeals, the superior courts and the district courts. The superior and district courts are trial courts. Superior courts are courts of general jurisdiction, while district courts hear only certain types of cases, including misdemeanor criminal cases and civil cases valued up to $100,000.
The Supreme Court and the Court of Appeals are appellate courts. The Court of Appeals is required to hear appeals from certain lower-court decisions, including those regarding criminal prosecutions, juvenile delinquency, and habeas corpus. The Supreme Court hears civil appeals and may in its discretion hear criminal appeals.
State politics
Although in its early years of statehood Alaska was a Democratic state, since the early 1970s it has been characterized as Republican-leaning. Local political communities have often worked on issues related to land use development, fishing, tourism, and individual rights. Alaska Natives, while organized in and around their communities, have been active within the Native corporations. These have been given ownership over large tracts of land, which require stewardship.
Alaska was formerly the only state in which possession of one ounce or less of marijuana in one's home was completely legal under state law, though the federal law remains in force.
The state has an independence movement favoring a vote on secession from the United States, with the Alaskan Independence Party. | [
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624_30 | Six Republicans and four Democrats have served as governor of Alaska. In addition, Republican governor Wally Hickel was elected to the office for a second term in 1990 after leaving the Republican party and briefly joining the Alaskan Independence Party ticket just long enough to be reelected. He officially rejoined the Republican party in 1994.
Alaska's voter initiative making marijuana legal took effect on February 24, 2015, placing Alaska alongside Colorado and Washington as the first three U.S. states where recreational marijuana is legal. The new law means people over 21 can consume small amounts of cannabis. The first legal marijuana store opened in Valdez in October 2016.
Voter registration
Taxes
To finance state government operations, Alaska depends primarily on petroleum revenues and federal subsidies. This allows it to have the lowest individual tax burden in the United States. It is one of five states with no sales tax, one of seven states with no individual income tax, and—along with New Hampshire—one of two that has neither. The Department of Revenue Tax Division reports regularly on the state's revenue sources. The Department also issues an annual summary of its operations, including new state laws that directly affect the tax division. In 2014, the Tax Foundation ranked Alaska as having the fourth most "business friendly" tax policy, behind only Wyoming, South Dakota, and Nevada.
While Alaska has no state sales tax, 89 municipalities collect a local sales tax, from 1.0 to 7.5%, typically 3–5%. Other local taxes levied include raw fish taxes, hotel, motel, and bed-and-breakfast 'bed' taxes, severance taxes, liquor and tobacco taxes, gaming (pull tabs) taxes, tire taxes and fuel transfer taxes. A part of the revenue collected from certain state taxes and license fees (such as petroleum, aviation motor fuel, telephone cooperative) is shared with municipalities in Alaska.
The fall in oil prices after the fracking boom in the early 2010s has decimated Alaska's state treasury, which has historically received about 85 percent of its revenue from taxes and fees imposed on oil and gas companies. The state government has had to drastically reduce its budget, and has brought its budget shortfall from over $2 billion in 2016 to under $500 million by 2018. In 2020, Alaska's state government budget was $4.8 billion, while projected government revenues were only $4.5 billion. | [
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] |
624_31 | Federal politics
Alaska regularly supports Republicans in presidential elections and has done so since statehood. Republicans have won the state's electoral college votes in all but one election that it has participated in (1964). No state has voted for a Democratic presidential candidate fewer times. Alaska was carried by Democratic nominee Lyndon B. Johnson during his landslide election in 1964, while the 1960 and 1968 elections were close. Since 1972, however, Republicans have carried the state by large margins. In 2008, Republican John McCain defeated Democrat Barack Obama in Alaska, 59.49% to 37.83%. McCain's running mate was Sarah Palin, the state's governor and the first Alaskan on a major party ticket. Obama lost Alaska again in 2012, but he captured 40% of the state's vote in that election, making him the first Democrat to do so since 1968.
The Alaska Bush, central Juneau, midtown and downtown Anchorage, and the areas surrounding the University of Alaska Fairbanks campus and Ester have been strongholds of the Democratic Party. The Matanuska-Susitna Borough, the majority of Fairbanks (including North Pole and the military base), and South Anchorage typically have the strongest Republican showing.
Elections
In a 2020 study, Alaska was ranked as the 15th hardest state for citizens to vote in.
In the 2020 election cycle, Alaskan voters approved Ballot Measure 2. The measure passed by a margin of 1.1%, or about 4,000 votes. The measure requires campaigns to disclose the original source and any intermediaries for campaign contributions over $2,000. The measure establishes non-partisan blanket primaries for statewide elections (like in Washington state and California) and ranked-choice voting (like in Maine). Alaska is the third state with jungle primaries for all statewide races, the second state with ranked voting, and the only state with both.
The first race to use the new system of elections will be the 2022 Senate election in which Lisa Murkowski will run for re-election.
See also
Index of Alaska-related articles
Outline of Alaska
Notes
References
External links | [
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] |
624_32 | Notes
References
External links
Alaska's Digital Archives
Alaska Inter-Tribal Council
Who Owns/Manages Alaska? (map)
Carl J. Sacarlasen Diary Extracts at Dartmouth College Library
M.E. Diemer Alaska Photographs at Dartmouth College Library
Alfred Hulse Brooks Photographs and Papers. Yale Collection of Western Americana, Beinecke Rare Book and Manuscript Library.
U.S. federal government
Alaska State Guide from the Library of Congress
Energy & Environmental Data for Alaska
USGS real-time, geographic, and other scientific resources of Alaska
US Census Bureau
Alaska State Facts
Alaska Statehood Subject Guide from the Eisenhower Presidential Library
Alaska Statehood documents, Dwight D. Eisenhower Presidential Library
Alaska state government
State of Alaska website
Alaska State Databases
Alaska Department of Natural Resources, Recorder's Office
Arctic Ocean
Former Russian colonies
States and territories established in 1959
States of the United States
States of the West Coast of the United States
U.S. states with multiple time zones
1959 establishments in the United States
Western United States
Northern America
Enclaves and exclaves
Beringia | [
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627_0 | Agriculture is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled people to live in cities. The history of agriculture began thousands of years ago. After gathering wild grains beginning at least 105,000 years ago, nascent farmers began to plant them around 11,500 years ago. Pigs, sheep, and cattle were domesticated over 10,000 years ago. Plants were independently cultivated in at least 11 regions of the world. Industrial agriculture based on large-scale monoculture in the twentieth century came to dominate agricultural output, though about 2 billion people still depended on subsistence agriculture.
The major agricultural products can be broadly grouped into foods, fibers, fuels and raw materials (such as rubber). Food classes include cereals (grains), vegetables, fruits, oils, meat, milk, eggs and fungi. Over one-third of the world's workers are employed in agriculture, second only to the service sector, although in recent decades, the global trend of a decreasing number of agricultural workers continues, especially in developing countries where smallholding is being overtaken by industrial agriculture and mechanization that brings an enormous crop yield increase.
Modern agronomy, plant breeding, agrochemicals such as pesticides and fertilizers, and technological developments have sharply increased crop yields, but causing ecological and environmental damage. Selective breeding and modern practices in animal husbandry have similarly increased the output of meat, but have raised concerns about animal welfare and environmental damage. Environmental issues include contributions to global warming, depletion of aquifers, deforestation, antibiotic resistance, and growth hormones in industrial meat production. Agriculture is both a cause of and sensitive to environmental degradation, such as biodiversity loss, desertification, soil degradation and global warming, all of which can cause decreases in crop yield. Genetically modified organisms are widely used, although some are banned in certain countries.
Etymology and scope | [
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627_1 | Etymology and scope
The word agriculture is a late Middle English adaptation of Latin , from 'field' and 'cultivation' or 'growing'. While agriculture usually refers to human activities, certain species of ant, termite and beetle have been cultivating crops for up to 60 million years. Agriculture is defined with varying scopes, in its broadest sense using natural resources to "produce commodities which maintain life, including food, fiber, forest products, horticultural crops, and their related services". Thus defined, it includes arable farming, horticulture, animal husbandry and forestry, but horticulture and forestry are in practice often excluded.
History
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] |
627_2 | History
Origins
The development of agriculture enabled the human population to grow many times larger than could be sustained by hunting and gathering. Agriculture began independently in different parts of the globe, and included a diverse range of taxa, in at least 11 separate centres of origin. Wild grains were collected and eaten from at least 105,000 years ago. From around 11,500 years ago, the eight Neolithic founder crops, emmer and einkorn wheat, hulled barley, peas, lentils, bitter vetch, chick peas and flax were cultivated in the Levant. Rice was domesticated in China between 11,500 and 6,200 BC with the earliest known cultivation from 5,700 BC, followed by mung, soy and azuki beans. Sheep were domesticated in Mesopotamia between 13,000 and 11,000 years ago. Cattle were domesticated from the wild aurochs in the areas of modern Turkey and Pakistan some 10,500 years ago. Pig production emerged in Eurasia, including Europe, East Asia and Southwest Asia, where wild boar were first domesticated about 10,500 years ago. In the Andes of South America, the potato was domesticated between 10,000 and 7,000 years ago, along with beans, coca, llamas, alpacas, and guinea pigs. Sugarcane and some root vegetables were domesticated in New Guinea around 9,000 years ago. Sorghum was domesticated in the Sahel region of Africa by 7,000 years ago. Cotton was domesticated in Peru by 5,600 years ago, and was independently domesticated in Eurasia. In Mesoamerica, wild teosinte was bred into maize by 6,000 years ago.
Scholars have offered multiple hypotheses to explain the historical origins of agriculture. Studies of the transition from hunter-gatherer to agricultural societies indicate an initial period of intensification and increasing sedentism; examples are the Natufian culture in the Levant, and the Early Chinese Neolithic in China. Then, wild stands that had previously been harvested started to be planted, and gradually came to be domesticated. | [
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627_3 | Civilizations
In Eurasia, the Sumerians started to live in villages from about 8,000 BC, relying on the Tigris and Euphrates rivers and a canal system for irrigation. Ploughs appear in pictographs around 3,000 BC; seed-ploughs around 2,300 BC. Farmers grew wheat, barley, vegetables such as lentils and onions, and fruits including dates, grapes, and figs. Ancient Egyptian agriculture relied on the Nile River and its seasonal flooding. Farming started in the predynastic period at the end of the Paleolithic, after 10,000 BC. Staple food crops were grains such as wheat and barley, alongside industrial crops such as flax and papyrus. In India, wheat, barley and jujube were domesticated by 9,000 BC, soon followed by sheep and goats. Cattle, sheep and goats were domesticated in Mehrgarh culture by 8,000–6,000 BC. Cotton was cultivated by the 5th–4th millennium BC. Archeological evidence indicates an animal-drawn plough from 2,500 BC in the Indus Valley Civilisation.
In China, from the 5th century BC there was a nationwide granary system and widespread silk farming. Water-powered grain mills were in use by the 1st century BC, followed by irrigation. By the late 2nd century, heavy ploughs had been developed with iron ploughshares and mouldboards. These spread westwards across Eurasia. Asian rice was domesticated 8,200–13,500 years ago – depending on the molecular clock estimate that is used – on the Pearl River in southern China with a single genetic origin from the wild rice Oryza rufipogon. In Greece and Rome, the major cereals were wheat, emmer, and barley, alongside vegetables including peas, beans, and olives. Sheep and goats were kept mainly for dairy products. | [
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627_4 | In the Americas, crops domesticated in Mesoamerica (apart from teosinte) include squash, beans, and cacao. Cocoa was being domesticated by the Mayo Chinchipe of the upper Amazon around 3,000 BC.
The turkey was probably domesticated in Mexico or the American Southwest. The Aztecs developed irrigation systems, formed terraced hillsides, fertilized their soil, and developed chinampas or artificial islands. The Mayas used extensive canal and raised field systems to farm swampland from 400 BC. Coca was domesticated in the Andes, as were the peanut, tomato, tobacco, and pineapple. Cotton was domesticated in Peru by 3,600 BC. Animals including llamas, alpacas, and guinea pigs were domesticated there. In North America, the indigenous people of the East domesticated crops such as sunflower, tobacco, squash and Chenopodium. Wild foods including wild rice and maple sugar were harvested. The domesticated strawberry is a hybrid of a Chilean and a North American species, developed by breeding in Europe and North America. The indigenous people of the Southwest and the Pacific Northwest practiced forest gardening and fire-stick farming. The natives controlled fire on a regional scale to create a low-intensity fire ecology that sustained a low-density agriculture in loose rotation; a sort of "wild" permaculture. A system of companion planting called the Three Sisters was developed in North America. The three crops were winter squash, maize, and climbing beans.
Indigenous Australians, long supposed to have been nomadic hunter-gatherers, practised systematic burning, possibly to enhance natural productivity in fire-stick farming. Scholars have pointed out that hunter-gatherers need a productive environment to support gathering without cultivation. Because the forests of New Guinea have few food plants, early humans may have used "selective burning" to increase the productivity of the wild karuka fruit trees to support the hunter-gatherer way of life.
The Gunditjmara and other groups developed eel farming and fish trapping systems from some 5,000 years ago. There is evidence of 'intensification' across the whole continent over that period. In two regions of Australia, the central west coast and eastern central, early farmers cultivated yams, native millet, and bush onions, possibly in permanent settlements.
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] |
627_5 | Revolution
In the Middle Ages, both in Europe and in the Islamic world, agriculture transformed with improved techniques and the diffusion of crop plants, including the introduction of sugar, rice, cotton and fruit trees (such as the orange) to Europe by way of Al-Andalus.
After 1492 the Columbian exchange brought New World crops such as maize, potatoes, tomatoes, sweet potatoes and manioc to Europe, and Old World crops such as wheat, barley, rice and turnips, and livestock (including horses, cattle, sheep and goats) to the Americas.
Irrigation, crop rotation, and fertilizers advanced from the 17th century with the British Agricultural Revolution, allowing global population to rise significantly. Since 1900 agriculture in developed nations, and to a lesser extent in the developing world, has seen large rises in productivity as mechanization replaces human labor, and assisted by synthetic fertilizers, pesticides, and selective breeding. The Haber-Bosch method allowed the synthesis of ammonium nitrate fertilizer on an industrial scale, greatly increasing crop yields and sustaining a further increase in global population. Modern agriculture has raised or encountered ecological, political, and economic issues including water pollution, biofuels, genetically modified organisms, tariffs and farm subsidies, leading to alternative approaches such as the organic movement.
Types
Pastoralism involves managing domesticated animals. In nomadic pastoralism, herds of livestock are moved from place to place in search of pasture, fodder, and water. This type of farming is practised in arid and semi-arid regions of Sahara, Central Asia and some parts of India.
In shifting cultivation, a small area of forest is cleared by cutting and burning the trees. The cleared land is used for growing crops for a few years until the soil becomes too infertile, and the area is abandoned. Another patch of land is selected and the process is repeated. This type of farming is practiced mainly in areas with abundant rainfall where the forest regenerates quickly. This practice is used in Northeast India, Southeast Asia, and the Amazon Basin.
Subsistence farming is practiced to satisfy family or local needs alone, with little left over for transport elsewhere. It is intensively practiced in Monsoon Asia and South-East Asia. An estimated 2.5 billion subsistence farmers worked in 2018, cultivating about 60% of the earth's arable land. | [
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627_6 | Intensive farming is cultivation to maximise productivity, with a low fallow ratio and a high use of inputs (water, fertilizer, pesticide and automation). It is practiced mainly in developed countries.
Contemporary agriculture
Status
From the twentieth century, intensive agriculture increased productivity. It substituted synthetic fertilizers and pesticides for labour, but caused increased water pollution, and often involved farm subsidies. In recent years there has been a backlash against the environmental effects of conventional agriculture, resulting in the organic, regenerative, and sustainable agriculture movements. One of the major forces behind this movement has been the European Union, which first certified organic food in 1991 and began reform of its Common Agricultural Policy (CAP) in 2005 to phase out commodity-linked farm subsidies, also known as decoupling. The growth of organic farming has renewed research in alternative technologies such as integrated pest management, selective breeding, and controlled-environment agriculture. Recent mainstream technological developments include genetically modified food. Demand for non-food biofuel crops, development of former farm lands, rising transportation costs, climate change, growing consumer demand in China and India, and population growth, are threatening food security in many parts of the world. The International Fund for Agricultural Development posits that an increase in smallholder agriculture may be part of the solution to concerns about food prices and overall food security, given the favorable experience of Vietnam. Soil degradation and diseases such as stem rust are major concerns globally; approximately 40% of the world's agricultural land is seriously degraded. By 2015, the agricultural output of China was the largest in the world, followed by the European Union, India and the United States. Economists measure the total factor productivity of agriculture and by this measure agriculture in the United States is roughly 1.7 times more productive than it was in 1948.
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627_7 | Workforce
Following the three-sector theory, the number of people employed in agriculture and other primary activities (such as fishing) can be more than 80% in the least developed countries, and less than 2% in the most highly developed countries. Since the Industrial Revolution, many countries have made the transition to developed economies, and the proportion of people working in agriculture has steadily fallen. During the 16th century in Europe, for example, between 55 and 75% of the population was engaged in agriculture; by the 19th century, this had dropped to between 35 and 65%. In the same countries today, the figure is less than 10%.
At the start of the 21st century, some one billion people, or over 1/3 of the available work force, were employed in agriculture. It constitutes approximately 70% of the global employment of children, and in many countries employs the largest percentage of women of any industry. The service sector overtook the agricultural sector as the largest global employer in 2007.
Safety
Agriculture, specifically farming, remains a hazardous industry, and farmers worldwide remain at high risk of work-related injuries, lung disease, noise-induced hearing loss, skin diseases, as well as certain cancers related to chemical use and prolonged sun exposure. On industrialized farms, injuries frequently involve the use of agricultural machinery, and a common cause of fatal agricultural injuries in developed countries is tractor rollovers. Pesticides and other chemicals used in farming can be hazardous to worker health, and workers exposed to pesticides may experience illness or have children with birth defects. As an industry in which families commonly share in work and live on the farm itself, entire families can be at risk for injuries, illness, and death. Ages 0–6 May be an especially vulnerable population in agriculture; common causes of fatal injuries among young farm workers include drowning, machinery and motor accidents, including with all-terrain vehicles.
The International Labour Organization considers agriculture "one of the most hazardous of all economic sectors". It estimates that the annual work-related death toll among agricultural employees is at least 170,000, twice the average rate of other jobs. In addition, incidences of death, injury and illness related to agricultural activities often go unreported. The organization has developed the Safety and Health in Agriculture Convention, 2001, which covers the range of risks in the agriculture occupation, the prevention of these risks and the role that individuals and organizations engaged in agriculture should play. | [
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627_8 | In the United States, agriculture has been identified by the National Institute for Occupational Safety and Health as a priority industry sector in the National Occupational Research Agenda to identify and provide intervention strategies for occupational health and safety issues.
In the European Union, the European Agency for Safety and Health at Work has issued guidelines on implementing health and safety directives in agriculture, livestock farming, horticulture, and forestry. The Agricultural Safety and Health Council of America (ASHCA) also holds a yearly summit to discuss safety.
Production
Overall production varies by country as listed.
Crop cultivation systems
Cropping systems vary among farms depending on the available resources and constraints; geography and climate of the farm; government policy; economic, social and political pressures; and the philosophy and culture of the farmer.
Shifting cultivation (or slash and burn) is a system in which forests are burnt, releasing nutrients to support cultivation of annual and then perennial crops for a period of several years. Then the plot is left fallow to regrow forest, and the farmer moves to a new plot, returning after many more years (10–20). This fallow period is shortened if population density grows, requiring the input of nutrients (fertilizer or manure) and some manual pest control. Annual cultivation is the next phase of intensity in which there is no fallow period. This requires even greater nutrient and pest control inputs.
Further industrialization led to the use of monocultures, when one cultivar is planted on a large acreage. Because of the low biodiversity, nutrient use is uniform and pests tend to build up, necessitating the greater use of pesticides and fertilizers. Multiple cropping, in which several crops are grown sequentially in one year, and intercropping, when several crops are grown at the same time, are other kinds of annual cropping systems known as polycultures.
In subtropical and arid environments, the timing and extent of agriculture may be limited by rainfall, either not allowing multiple annual crops in a year, or requiring irrigation. In all of these environments perennial crops are grown (coffee, chocolate) and systems are practiced such as agroforestry. In temperate environments, where ecosystems were predominantly grassland or prairie, highly productive annual farming is the dominant agricultural system. | [
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627_9 | Important categories of food crops include cereals, legumes, forage, fruits and vegetables. Natural fibers include cotton, wool, hemp, silk and flax. Specific crops are cultivated in distinct growing regions throughout the world. Production is listed in millions of metric tons, based on FAO estimates.
Livestock production systems
Animal husbandry is the breeding and raising of animals for meat, milk, eggs, or wool, and for work and transport. Working animals, including horses, mules, oxen, water buffalo, camels, llamas, alpacas, donkeys, and dogs, have for centuries been used to help cultivate fields, harvest crops, wrangle other animals, and transport farm products to buyers.
Livestock production systems can be defined based on feed source, as grassland-based, mixed, and landless. , 30% of Earth's ice- and water-free area was used for producing livestock, with the sector employing approximately 1.3 billion people. Between the 1960s and the 2000s, there was a significant increase in livestock production, both by numbers and by carcass weight, especially among beef, pigs and chickens, the latter of which had production increased by almost a factor of 10. Non-meat animals, such as milk cows and egg-producing chickens, also showed significant production increases. Global cattle, sheep and goat populations are expected to continue to increase sharply through 2050. Aquaculture or fish farming, the production of fish for human consumption in confined operations, is one of the fastest growing sectors of food production, growing at an average of 9% a year between 1975 and 2007.
During the second half of the 20th century, producers using selective breeding focused on creating livestock breeds and crossbreeds that increased production, while mostly disregarding the need to preserve genetic diversity. This trend has led to a significant decrease in genetic diversity and resources among livestock breeds, leading to a corresponding decrease in disease resistance and local adaptations previously found among traditional breeds. | [
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627_10 | Grassland based livestock production relies upon plant material such as shrubland, rangeland, and pastures for feeding ruminant animals. Outside nutrient inputs may be used, however manure is returned directly to the grassland as a major nutrient source. This system is particularly important in areas where crop production is not feasible because of climate or soil, representing 30–40 million pastoralists. Mixed production systems use grassland, fodder crops and grain feed crops as feed for ruminant and monogastric (one stomach; mainly chickens and pigs) livestock. Manure is typically recycled in mixed systems as a fertilizer for crops.
Landless systems rely upon feed from outside the farm, representing the de-linking of crop and livestock production found more prevalently in Organisation for Economic Co-operation and Development member countries. Synthetic fertilizers are more heavily relied upon for crop production and manure use becomes a challenge as well as a source for pollution. Industrialized countries use these operations to produce much of the global supplies of poultry and pork. Scientists estimate that 75% of the growth in livestock production between 2003 and 2030 will be in confined animal feeding operations, sometimes called factory farming. Much of this growth is happening in developing countries in Asia, with much smaller amounts of growth in Africa. Some of the practices used in commercial livestock production, including the usage of growth hormones, are controversial.
Production practices
Tillage is the practice of breaking up the soil with tools such as the plow or harrow to prepare for planting, for nutrient incorporation, or for pest control. Tillage varies in intensity from conventional to no-till. It may improve productivity by warming the soil, incorporating fertilizer and controlling weeds, but also renders soil more prone to erosion, triggers the decomposition of organic matter releasing CO2, and reduces the abundance and diversity of soil organisms.
Pest control includes the management of weeds, insects, mites, and diseases. Chemical (pesticides), biological (biocontrol), mechanical (tillage), and cultural practices are used. Cultural practices include crop rotation, culling, cover crops, intercropping, composting, avoidance, and resistance. Integrated pest management attempts to use all of these methods to keep pest populations below the number which would cause economic loss, and recommends pesticides as a last resort. | [
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627_11 | Nutrient management includes both the source of nutrient inputs for crop and livestock production, and the method of use of manure produced by livestock. Nutrient inputs can be chemical inorganic fertilizers, manure, green manure, compost and minerals. Crop nutrient use may also be managed using cultural techniques such as crop rotation or a fallow period. Manure is used either by holding livestock where the feed crop is growing, such as in managed intensive rotational grazing, or by spreading either dry or liquid formulations of manure on cropland or pastures.
Water management is needed where rainfall is insufficient or variable, which occurs to some degree in most regions of the world. Some farmers use irrigation to supplement rainfall. In other areas such as the Great Plains in the U.S. and Canada, farmers use a fallow year to conserve soil moisture to use for growing a crop in the following year. Agriculture represents 70% of freshwater use worldwide.
According to a report by the International Food Policy Research Institute, agricultural technologies will have the greatest impact on food production if adopted in combination with each other; using a model that assessed how eleven technologies could impact agricultural productivity, food security and trade by 2050, the International Food Policy Research Institute found that the number of people at risk from hunger could be reduced by as much as 40% and food prices could be reduced by almost half.
Payment for ecosystem services is a method of providing additional incentives to encourage farmers to conserve some aspects of the environment. Measures might include paying for reforestation upstream of a city, to improve the supply of fresh water.
Crop alteration and biotechnology
Plant breeding
Crop alteration has been practiced by humankind for thousands of years, since the beginning of civilization. Altering crops through breeding practices changes the genetic make-up of a plant to develop crops with more beneficial characteristics for humans, for example, larger fruits or seeds, drought-tolerance, or resistance to pests. Significant advances in plant breeding ensued after the work of geneticist Gregor Mendel. His work on dominant and recessive alleles, although initially largely ignored for almost 50 years, gave plant breeders a better understanding of genetics and breeding techniques. Crop breeding includes techniques such as plant selection with desirable traits, self-pollination and cross-pollination, and molecular techniques that genetically modify the organism. | [
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627_12 | Domestication of plants has, over the centuries increased yield, improved disease resistance and drought tolerance, eased harvest and improved the taste and nutritional value of crop plants. Careful selection and breeding have had enormous effects on the characteristics of crop plants. Plant selection and breeding in the 1920s and 1930s improved pasture (grasses and clover) in New Zealand. Extensive X-ray and ultraviolet induced mutagenesis efforts (i.e. primitive genetic engineering) during the 1950s produced the modern commercial varieties of grains such as wheat, corn (maize) and barley.
The Green Revolution popularized the use of conventional hybridization to sharply increase yield by creating "high-yielding varieties". For example, average yields of corn (maize) in the US have increased from around 2.5 tons per hectare (t/ha) (40 bushels per acre) in 1900 to about 9.4 t/ha (150 bushels per acre) in 2001. Similarly, worldwide average wheat yields have increased from less than 1 t/ha in 1900 to more than 2.5 t/ha in 1990. South American average wheat yields are around 2 t/ha, African under 1 t/ha, and Egypt and Arabia up to 3.5 to 4 t/ha with irrigation. In contrast, the average wheat yield in countries such as France is over 8 t/ha. Variations in yields are due mainly to variation in climate, genetics, and the level of intensive farming techniques (use of fertilizers, chemical pest control, growth control to avoid lodging).
Genetic engineering
Genetically modified organisms (GMO) are organisms whose genetic material has been altered by genetic engineering techniques generally known as recombinant DNA technology. Genetic engineering has expanded the genes available to breeders to use in creating desired germlines for new crops. Increased durability, nutritional content, insect and virus resistance and herbicide tolerance are a few of the attributes bred into crops through genetic engineering. For some, GMO crops cause food safety and food labeling concerns. Numerous countries have placed restrictions on the production, import or use of GMO foods and crops. Currently a global treaty, the Biosafety Protocol, regulates the trade of GMOs. There is ongoing discussion regarding the labeling of foods made from GMOs, and while the EU currently requires all GMO foods to be labeled, the US does not. | [
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627_13 | Herbicide-resistant seed has a gene implanted into its genome that allows the plants to tolerate exposure to herbicides, including glyphosate. These seeds allow the farmer to grow a crop that can be sprayed with herbicides to control weeds without harming the resistant crop. Herbicide-tolerant crops are used by farmers worldwide. With the increasing use of herbicide-tolerant crops, comes an increase in the use of glyphosate-based herbicide sprays. In some areas glyphosate resistant weeds have developed, causing farmers to switch to other herbicides. Some studies also link widespread glyphosate usage to iron deficiencies in some crops, which is both a crop production and a nutritional quality concern, with potential economic and health implications.
Other GMO crops used by growers include insect-resistant crops, which have a gene from the soil bacterium Bacillus thuringiensis (Bt), which produces a toxin specific to insects. These crops resist damage by insects. Some believe that similar or better pest-resistance traits can be acquired through traditional breeding practices, and resistance to various pests can be gained through hybridization or cross-pollination with wild species. In some cases, wild species are the primary source of resistance traits; some tomato cultivars that have gained resistance to at least 19 diseases did so through crossing with wild populations of tomatoes.
Environmental impact
Effects and costs
Agriculture is both a cause of and sensitive to environmental degradation, such as biodiversity loss, desertification, soil degradation and global warming, which cause decrease in crop yield. Agriculture is one of the most important drivers of environmental pressures, particularly habitat change, climate change, water use and toxic emissions. Agriculture is the main source of toxins released into the environment, including insecticides, especially those used on cotton. The 2011 UNEP Green Economy report stated that agricultural operations produced some 13 per cent of anthropogenic global greenhouse gas emissions. This includes gases from the use of inorganic fertilizers, agro-chemical pesticides, and herbicides, as well as fossil fuel-energy inputs. | [
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627_14 | Agriculture imposes multiple external costs upon society through effects such as pesticide damage to nature (especially herbicides and insecticides), nutrient runoff, excessive water usage, and loss of natural environment. A 2000 assessment of agriculture in the UK determined total external costs for 1996 of £2,343 million, or £208 per hectare. A 2005 analysis of these costs in the US concluded that cropland imposes approximately $5 to $16 billion ($30 to $96 per hectare), while livestock production imposes $714 million. Both studies, which focused solely on the fiscal impacts, concluded that more should be done to internalize external costs. Neither included subsidies in their analysis, but they noted that subsidies also influence the cost of agriculture to society.
Agriculture seeks to increase yield and to reduce costs. Yield increases with inputs such as fertilisers and removal of pathogens, predators, and competitors (such as weeds). Costs decrease with increasing scale of farm units, such as making fields larger; this means removing hedges, ditches and other areas of habitat. Pesticides kill insects, plants and fungi. These and other measures have cut biodiversity to very low levels on intensively farmed land. Effective yields fall with on-farm losses, which may be caused by poor production practices during harvesting, handling, and storage.
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627_15 | Livestock issues
A senior UN official, Henning Steinfeld, said that "Livestock are one of the most significant contributors to today's most serious environmental problems". Livestock production occupies 70% of all land used for agriculture, or 30% of the land surface of the planet. It is one of the largest sources of greenhouse gases, responsible for 18% of the world's greenhouse gas emissions as measured in CO2 equivalents. By comparison, all transportation emits 13.5% of the CO2. It produces 65% of human-related nitrous oxide (which has 296 times the global warming potential of CO2) and 37% of all human-induced methane (which is 23 times as warming as CO2.) It also generates 64% of the ammonia emission. Livestock expansion is cited as a key factor driving deforestation; in the Amazon basin 70% of previously forested area is now occupied by pastures and the remainder used for feedcrops. Through deforestation and land degradation, livestock is also driving reductions in biodiversity. Furthermore, the UNEP states that "methane emissions from global livestock are projected to increase by 60 per cent by 2030 under current practices and consumption patterns."
Land and water issues
Land transformation, the use of land to yield goods and services, is the most substantial way humans alter the Earth's ecosystems, and is the driving force causing biodiversity loss. Estimates of the amount of land transformed by humans vary from 39 to 50%. Land degradation, the long-term decline in ecosystem function and productivity, is estimated to be occurring on 24% of land worldwide, with cropland overrepresented. Land management is the driving factor behind degradation; 1.5 billion people rely upon the degrading land. Degradation can be through deforestation, desertification, soil erosion, mineral depletion, acidification, or salinization. | [
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627_16 | Eutrophication, excessive nutrient enrichment in aquatic ecosystems resulting in algal blooms and anoxia, leads to fish kills, loss of biodiversity, and renders water unfit for drinking and other industrial uses. Excessive fertilization and manure application to cropland, as well as high livestock stocking densities cause nutrient (mainly nitrogen and phosphorus) runoff and leaching from agricultural land. These nutrients are major nonpoint pollutants contributing to eutrophication of aquatic ecosystems and pollution of groundwater, with harmful effects on human populations. Fertilisers also reduce terrestrial biodiversity by increasing competition for light, favouring those species that are able to benefit from the added nutrients.
Agriculture accounts for 70 percent of withdrawals of freshwater resources. Agriculture is a major draw on water from aquifers, and currently draws from those underground water sources at an unsustainable rate. It is long known that aquifers in areas as diverse as northern China, the Upper Ganges and the western US are being depleted, and new research extends these problems to aquifers in Iran, Mexico and Saudi Arabia. Increasing pressure is being placed on water resources by industry and urban areas, meaning that water scarcity is increasing and agriculture is facing the challenge of producing more food for the world's growing population with reduced water resources. Agricultural water usage can also cause major environmental problems, including the destruction of natural wetlands, the spread of water-borne diseases, and land degradation through salinization and waterlogging, when irrigation is performed incorrectly.
Pesticides
Pesticide use has increased since 1950 to 2.5million short tons annually worldwide, yet crop loss from pests has remained relatively constant. The World Health Organization estimated in 1992 that three million pesticide poisonings occur annually, causing 220,000 deaths. Pesticides select for pesticide resistance in the pest population, leading to a condition termed the "pesticide treadmill" in which pest resistance warrants the development of a new pesticide. | [
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627_17 | An alternative argument is that the way to "save the environment" and prevent famine is by using pesticides and intensive high yield farming, a view exemplified by a quote heading the Center for Global Food Issues website: 'Growing more per acre leaves more land for nature'. However, critics argue that a trade-off between the environment and a need for food is not inevitable, and that pesticides simply replace good agronomic practices such as crop rotation. The Push–pull agricultural pest management technique involves intercropping, using plant aromas to repel pests from crops (push) and to lure them to a place from which they can then be removed (pull).
Climate change
Climate change and agriculture are interrelated on a global scale. Global warming affects agriculture through changes in average temperatures, rainfall, and weather extremes (like storms and heat waves); changes in pests and diseases; changes in atmospheric carbon dioxide and ground-level ozone concentrations; changes in the nutritional quality of some foods; and changes in sea level. Global warming is already affecting agriculture, with effects unevenly distributed across the world. Future climate change will probably negatively affect crop production in low latitude countries, while effects in northern latitudes may be positive or negative. Global warming will probably increase the risk of food insecurity for some vulnerable groups, such as the poor.
Animal husbandry is also responsible for greenhouse gas production of CO2 and a percentage of the world's methane, and future land infertility, and the displacement of wildlife. Agriculture contributes to climate change by anthropogenic emissions of greenhouse gases, and by the conversion of non-agricultural land such as forest for agricultural use. Agriculture, forestry and land-use change contributed around 20 to 25% to global annual emissions in 2010. A range of policies can reduce the risk of negative climate change impacts on agriculture, and greenhouse gas emissions from the agriculture sector.
Sustainability | [
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627_18 | Sustainability
Current farming methods have resulted in over-stretched water resources, high levels of erosion and reduced soil fertility. There is not enough water to continue farming using current practices; therefore how critical water, land, and ecosystem resources are used to boost crop yields must be reconsidered. A solution would be to give value to ecosystems, recognizing environmental and livelihood tradeoffs, and balancing the rights of a variety of users and interests. Inequities that result when such measures are adopted would need to be addressed, such as the reallocation of water from poor to rich, the clearing of land to make way for more productive farmland, or the preservation of a wetland system that limits fishing rights.
Technological advancements help provide farmers with tools and resources to make farming more sustainable. Technology permits innovations like conservation tillage, a farming process which helps prevent land loss to erosion, reduces water pollution, and enhances carbon sequestration. Other potential practices include conservation agriculture, agroforestry, improved grazing, avoided grassland conversion, and biochar. Current mono-crop farming practices in the United States preclude widespread adoption of sustainable practices, such as 2-3 crop rotations that incorporate grass or hay with annual crops, unless negative emission goals such as soil carbon sequestration become policy.
The International Food Policy Research Institute states that agricultural technologies will have the greatest impact on food production if adopted in combination with each other; using a model that assessed how eleven technologies could impact agricultural productivity, food security and trade by 2050, it found that the number of people at risk from hunger could be reduced by as much as 40% and food prices could be reduced by almost half. The food demand of Earth's projected population, with current climate change predictions, could be satisfied by improvement of agricultural methods, expansion of agricultural areas, and a sustainability-oriented consumer mindset.
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] |
627_19 | Energy dependence
Since the 1940s, agricultural productivity has increased dramatically, due largely to the increased use of energy-intensive mechanization, fertilizers and pesticides. The vast majority of this energy input comes from fossil fuel sources. Between the 1960s and the 1980s, the Green Revolution transformed agriculture around the globe, with world grain production increasing significantly (between 70% and 390% for wheat and 60% to 150% for rice, depending on geographic area) as world population doubled. Heavy reliance on petrochemicals has raised concerns that oil shortages could increase costs and reduce agricultural output.
Industrialized agriculture depends on fossil fuels in two fundamental ways: direct consumption on the farm and manufacture of inputs used on the farm. Direct consumption includes the use of lubricants and fuels to operate farm vehicles and machinery.
Indirect consumption includes the manufacture of fertilizers, pesticides, and farm machinery. In particular, the production of nitrogen fertilizer can account for over half of agricultural energy usage. Together, direct and indirect consumption by US farms accounts for about 2% of the nation's energy use. Direct and indirect energy consumption by U.S. farms peaked in 1979, and has since gradually declined. Food systems encompass not just agriculture but off-farm processing, packaging, transporting, marketing, consumption, and disposal of food and food-related items. Agriculture accounts for less than one-fifth of food system energy use in the US.
Disciplines
Agricultural economics | [
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] |
627_20 | Disciplines
Agricultural economics
Agricultural economics is economics as it relates to the "production, distribution and consumption of [agricultural] goods and services". Combining agricultural production with general theories of marketing and business as a discipline of study began in the late 1800s, and grew significantly through the 20th century. Although the study of agricultural economics is relatively recent, major trends in agriculture have significantly affected national and international economies throughout history, ranging from tenant farmers and sharecropping in the post-American Civil War Southern United States to the European feudal system of manorialism. In the United States, and elsewhere, food costs attributed to food processing, distribution, and agricultural marketing, sometimes referred to as the value chain, have risen while the costs attributed to farming have declined. This is related to the greater efficiency of farming, combined with the increased level of value addition (e.g. more highly processed products) provided by the supply chain. Market concentration has increased in the sector as well, and although the total effect of the increased market concentration is likely increased efficiency, the changes redistribute economic surplus from producers (farmers) and consumers, and may have negative implications for rural communities.
National government policies can significantly change the economic marketplace for agricultural products, in the form of taxation, subsidies, tariffs and other measures. Since at least the 1960s, a combination of trade restrictions, exchange rate policies and subsidies have affected farmers in both the developing and the developed world. In the 1980s, non-subsidized farmers in developing countries experienced adverse effects from national policies that created artificially low global prices for farm products. Between the mid-1980s and the early 2000s, several international agreements limited agricultural tariffs, subsidies and other trade restrictions. | [
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627_21 | However, , there was still a significant amount of policy-driven distortion in global agricultural product prices. The three agricultural products with the most trade distortion were sugar, milk and rice, mainly due to taxation. Among the oilseeds, sesame had the most taxation, but overall, feed grains and oilseeds had much lower levels of taxation than livestock products. Since the 1980s, policy-driven distortions have seen a greater decrease among livestock products than crops during the worldwide reforms in agricultural policy. Despite this progress, certain crops, such as cotton, still see subsidies in developed countries artificially deflating global prices, causing hardship in developing countries with non-subsidized farmers. Unprocessed commodities such as corn, soybeans, and cattle are generally graded to indicate quality, affecting the price the producer receives. Commodities are generally reported by production quantities, such as volume, number or weight.
Agricultural science
Agricultural science is a broad multidisciplinary field of biology that encompasses the parts of exact, natural, economic and social sciences used in the practice and understanding of agriculture. It covers topics such as agronomy, plant breeding and genetics, plant pathology, crop modelling, soil science, entomology, production techniques and improvement, study of pests and their management, and study of adverse environmental effects such as soil degradation, waste management, and bioremediation.
The scientific study of agriculture began in the 18th century, when Johann Friedrich Mayer conducted experiments on the use of gypsum (hydrated calcium sulphate) as a fertilizer. Research became more systematic when in 1843, John Lawes and Henry Gilbert began a set of long-term agronomy field experiments at Rothamsted Research Station in England; some of them, such as the Park Grass Experiment, are still running. In America, the Hatch Act of 1887 provided funding for what it was the first to call "agricultural science", driven by farmers' interest in fertilizers. In agricultural entomology, the USDA began to research biological control in 1881; it instituted its first large program in 1905, searching Europe and Japan for natural enemies of the gypsy moth and brown-tail moth, establishing parasitoids (such as solitary wasps) and predators of both pests in the USA.
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] |
627_22 | Policy
Agricultural policy is the set of government decisions and actions relating to domestic agriculture and imports of foreign agricultural products. Governments usually implement agricultural policies with the goal of achieving a specific outcome in the domestic agricultural product markets. Some overarching themes include risk management and adjustment (including policies related to climate change, food safety and natural disasters), economic stability (including policies related to taxes), natural resources and environmental sustainability (especially water policy), research and development, and market access for domestic commodities (including relations with global organizations and agreements with other countries). Agricultural policy can also touch on food quality, ensuring that the food supply is of a consistent and known quality, food security, ensuring that the food supply meets the population's needs, and conservation. Policy programs can range from financial programs, such as subsidies, to encouraging producers to enroll in voluntary quality assurance programs.
There are many influences on the creation of agricultural policy, including consumers, agribusiness, trade lobbies and other groups. Agribusiness interests hold a large amount of influence over policy making, in the form of lobbying and campaign contributions. Political action groups, including those interested in environmental issues and labor unions, also provide influence, as do lobbying organizations representing individual agricultural commodities. The Food and Agriculture Organization of the United Nations (FAO) leads international efforts to defeat hunger and provides a forum for the negotiation of global agricultural regulations and agreements. Samuel Jutzi, director of FAO's animal production and health division, states that lobbying by large corporations has stopped reforms that would improve human health and the environment. For example, proposals in 2010 for a voluntary code of conduct for the livestock industry that would have provided incentives for improving standards for health, and environmental regulations, such as the number of animals an area of land can support without long-term damage, were successfully defeated due to large food company pressure.
See also
Aeroponics
Agricultural aircraft
Agricultural engineering
Agricultural robot
Agroecology
Agrominerals
Building-integrated agriculture
Contract farming
Corporate farming
Crofting
Ecoagriculture
Hill farming
List of documentary films about agriculture
Pharming (genetics)
Remote sensing
Subsistence economy
Vertical farming
Vegetable farming
References
Cited sources
External links
Food and Agriculture Organization
United States Department of Agriculture
Agriculture material from the World Bank Group
Agriculture
Agronomy
Food industry | [
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628_0 | Aldous Leonard Huxley (26 July 1894 – 22 November 1963) was an English writer and philosopher. He wrote nearly 50 books—both novels and non-fiction works—as well as wide-ranging essays, narratives, and poems.
Born into the prominent Huxley family, he graduated from Balliol College, Oxford, with an undergraduate degree in English literature. Early in his career, he published short stories and poetry and edited the literary magazine Oxford Poetry, before going on to publish travel writing, satire, and screenplays. He spent the latter part of his life in the United States, living in Los Angeles from 1937 until his death. By the end of his life, Huxley was widely acknowledged as one of the foremost intellectuals of his time. He was nominated for the Nobel Prize in Literature nine times and was elected Companion of Literature by the Royal Society of Literature in 1962.
Huxley was a pacifist. He grew interested in philosophical mysticism and universalism, addressing these subjects with works such as The Perennial Philosophy (1945)—which illustrates commonalities between Western and Eastern mysticism—and The Doors of Perception (1954)—which interprets his own psychedelic experience with mescaline. In his most famous novel Brave New World (1932) and his final novel Island (1962), he presented his vision of dystopia and utopia, respectively.
Early life
Huxley was born in Godalming, Surrey, England, in 1894. He was the third son of the writer and schoolmaster Leonard Huxley, who edited The Cornhill Magazine, and his first wife, Julia Arnold, who founded Prior's Field School. Julia was the niece of poet and critic Matthew Arnold and the sister of Mrs. Humphry Ward. Julia named him Aldous after a character in one of her sister's novels. Aldous was the grandson of Thomas Henry Huxley, the zoologist, agnostic, and controversialist ("Darwin's Bulldog"). His brother Julian Huxley and half-brother Andrew Huxley also became outstanding biologists. Aldous had another brother, Noel Trevenen Huxley (1889–1914), who took his own life after a period of clinical depression. | [
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628_1 | As a child, Huxley's nickname was "Ogie", short for "Ogre". He was described by his brother, Julian, as someone who frequently "[contemplated] the strangeness of things". According to his cousin and contemporary, Gervas Huxley, he had an early interest in drawing.
Huxley's education began in his father's well-equipped botanical laboratory, after which he enrolled at Hillside School near Godalming. He was taught there by his own mother for several years until she became terminally ill. After Hillside he went on to Eton College. His mother died in 1908, when he was 14 (his father later remarried). He contracted the eye disease Keratitis punctata in 1911; this "left [him] practically blind for two to three years". This "ended his early dreams of becoming a doctor". In October 1913, Huxley entered Balliol College, Oxford, where he studied English literature. He volunteered for the British Army in January 1916, for the Great War; however, he was rejected on health grounds, being half-blind in one eye. His eyesight later partly recovered. He edited Oxford Poetry in 1916, and in June of that year graduated BA with first class honours. His brother Julian wrote:
Following his years at Balliol, Huxley, being financially indebted to his father, decided to find employment. He taught French for a year at Eton College, where Eric Blair (who was to take the pen name George Orwell) and Steven Runciman were among his pupils. He was mainly remembered as being an incompetent schoolmaster unable to keep order in class. Nevertheless, Blair and others spoke highly of his excellent command of language.
Huxley also worked for a time during the 1920s at Brunner and Mond, an advanced chemical plant in Billingham in County Durham, northeast England. According to the introduction to the latest edition of his science fiction novel Brave New World (1932), the experience he had there of "an ordered universe in a world of planless incoherence" was an important source for the novel.
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] |
628_2 | Career
Huxley completed his first (unpublished) novel at the age of 17 and began writing seriously in his early twenties, establishing himself as a successful writer and social satirist. His first published novels were social satires, Crome Yellow (1921), Antic Hay (1923), Those Barren Leaves (1925), and Point Counter Point (1928). Brave New World (1932) was his fifth novel and first dystopian work. In the 1920s, he was also a contributor to Vanity Fair and British Vogue magazines.
Contact with the Bloomsbury Set
During the First World War, Huxley spent much of his time at Garsington Manor near Oxford, home of Lady Ottoline Morrell, working as a farm labourer. While at the Manor, he met several Bloomsbury Group figures, including Bertrand Russell, Alfred North Whitehead, and Clive Bell. Later, in Crome Yellow (1921), he caricatured the Garsington lifestyle. Jobs were very scarce, but in 1919, John Middleton Murry was reorganising the Athenaeum and invited Huxley to join the staff. He accepted immediately, and quickly married the Belgian refugee Maria Nys (1899-1955), also at Garsington. They lived with their young son in Italy part of the time during the 1920s, where Huxley would visit his friend D. H. Lawrence. Following Lawrence's death in 1930, Huxley edited Lawrence's letters (1932). Very early in 1929, in London, Huxley met Gerald Heard, a brilliant writer and broadcaster, philosopher and interpreter of contemporary science.
Works of this period included important novels on the dehumanising aspects of scientific progress, most famously Brave New World, and on pacifist themes (for example, Eyeless in Gaza). In Brave New World, set in a dystopian London, Huxley portrays a society operating on the principles of mass production and Pavlovian conditioning. Huxley was strongly influenced by F. Matthias Alexander, and included him as a character in Eyeless in Gaza (1936).
Beginning in this period, Huxley began to write and edit non-fiction works on pacifist issues, including Ends and Means (1937), An Encyclopedia of Pacifism, and Pacifism and Philosophy, and was an active member of the Peace Pledge Union. | [
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] |
628_3 | Life in the United States
In 1937, Huxley moved to Hollywood with his wife Maria, son Matthew Huxley, and friend Gerald Heard. He lived in the U.S., mainly in southern California, until his death, and also for a time in Taos, New Mexico, where he wrote Ends and Means (published in 1937). The book contains tracts on war, religion, nationalism, and ethics.
Heard introduced Huxley to Vedanta (Upanishad-centered philosophy), meditation, and vegetarianism through the principle of ahimsa. In 1938, Huxley befriended Jiddu Krishnamurti, whose teachings he greatly admired. Huxley and Krishnamurti entered into an enduring exchange (sometimes edging on debate) over many years, with Krishnamurti representing the more rarefied, detached, ivory-tower perspective and Huxley, with his pragmatic concerns, the more socially and historically informed position. Huxley provided an introduction to Krishnamurti's quintessential statement, The First and Last Freedom (1954).
Huxley also became a Vedantist in the circle of Hindu Swami Prabhavananda, and introduced Christopher Isherwood to this circle. Not long afterwards, Huxley wrote his book on widely held spiritual values and ideas, The Perennial Philosophy, which discussed the teachings of renowned mystics of the world. Huxley's book affirmed a sensibility that insists there are realities beyond the generally accepted "five senses" and that there is genuine meaning for humans beyond both sensual satisfactions and sentimentalities.
Huxley became a close friend of Remsen Bird, president of Occidental College. He spent much time at the college, which is in the Eagle Rock neighbourhood of Los Angeles. The college appears as "Tarzana College" in his satirical novel After Many a Summer (1939). The novel won Huxley a British literary award, the 1939 James Tait Black Memorial Prize for fiction. Huxley also incorporated Bird into the novel. | [
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628_4 | During this period, Huxley earned a substantial income as a Hollywood screenwriter; Christopher Isherwood, in his autobiography My Guru and His Disciple, states that Huxley earned more than $3,000 per week (approximately $50,000 in 2020 dollars) as a screenwriter, and that he used much of it to transport Jewish and left-wing writer and artist refugees from Hitler's Germany to the US. In March 1938, Huxley's friend Anita Loos, a novelist and screenwriter, put him in touch with Metro-Goldwyn-Mayer (MGM), which hired him for Madame Curie which was originally to star Greta Garbo and be directed by George Cukor. (Eventually, the film was completed by MGM in 1943 with a different director and cast.) Huxley received screen credit for Pride and Prejudice (1940) and was paid for his work on a number of other films, including Jane Eyre (1944). He was commissioned by Walt Disney in 1945 to write a script based on Alice's Adventures in Wonderland and the biography of the story's author, Lewis Carroll. The script was not used, however.
Huxley wrote an introduction to the posthumous publication of J. D. Unwin's 1940 book Hopousia or The Sexual and Economic Foundations of a New Society.
On 21 October 1949, Huxley wrote to George Orwell, author of Nineteen Eighty-Four, congratulating him on "how fine and how profoundly important the book is". In his letter to Orwell, he predicted:
In 1953, Huxley and Maria applied for United States citizenship and presented themselves for examination. When Huxley refused to bear arms for the U.S. and would not state that his objections were based on religious ideals, the only excuse allowed under the McCarran Act, the judge had to adjourn the proceedings. He withdrew his application. Nevertheless, he remained in the U.S. In 1959, Huxley turned down an offer to be made a Knight Bachelor by the Macmillan government without putting forward a reason; his brother Julian had been knighted in 1958, while another brother Andrew would be knighted in 1974. | [
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628_5 | In the fall semester of 1960 Huxley was invited by Professor Huston Smith to be the Carnegie Visiting Professor of Humanities at the Massachusetts Institute of Technology (MIT). As part of the MIT centennial program of events organised by the Department of Humanities, Huxley presented a series of lectures titled, "What a Piece of Work is a Man" which concerned history, language, and art.
Late-in-life perspectives
Biographer Harold H. Watts wrote that Huxley's writings in the "final and extended period of his life" are "the work of a man who is meditating on the central problems of many modern men". Huxley had deeply felt apprehensions about the future the developed world might make for itself. From these, he made some warnings in his writings and talks. In a 1958 televised interview conducted by journalist Mike Wallace, Huxley outlined several major concerns: the difficulties and dangers of world overpopulation; the tendency towards distinctly hierarchical social organisation; the crucial importance of evaluating the use of technology in mass societies susceptible to persuasion; the tendency to promote modern politicians to a naive public as well-marketed commodities. In a December 1962 letter to brother Julian, summarizing a paper he had presented in Santa Barbara, he wrote, "What I said was that if we didn't pretty quickly start thinking of human problems in ecological terms rather than in terms of power politics we should very soon be in a bad way."
Huxley's engagement with Eastern wisdom traditions was entirely compatible with a strong appreciation of modern science. Biographer Milton Birnbaum wrote that Huxley "ended by embracing both science and Eastern religion". In his last book, Literature and Science, Huxley wrote that "The ethical and philosophical implications of modern science are more Buddhist than Christian...." In "A Philosopher's Visionary Prediction," published one month before he died, Huxley endorsed training in general semantics and "the nonverbal world of culturally uncontaminated consciousness," writing that "We must learn how to be mentally silent, we must cultivate the art of pure receptivity.... [T]he individual must learn to decondition himself, must be able to cut holes in the fence of verbalized symbols that hems him in." | [
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628_6 | Association with Vedanta
Beginning in 1939 and continuing until his death in 1963, Huxley had an extensive association with the Vedanta Society of Southern California, founded and headed by Swami Prabhavananda. Together with Gerald Heard, Christopher Isherwood and other followers, he was initiated by the Swami and was taught meditation and spiritual practices.
In 1944, Huxley wrote the introduction to the "Bhagavad Gita: The Song of God", translated by Swami Prabhavananda and Christopher Isherwood, which was published by the Vedanta Society of Southern California.
From 1941 until 1960, Huxley contributed 48 articles to Vedanta and the West, published by the society. He also served on the editorial board with Isherwood, Heard, and playwright John Van Druten from 1951 through 1962.
Huxley also occasionally lectured at the Hollywood and Santa Barbara Vedanta temples. Two of those lectures have been released on CD: Knowledge and Understanding and Who Are We? from 1955. Nonetheless, Huxley's agnosticism, together with his speculative propensity, made it difficult for him to fully embrace any form of institutionalised religion.
Psychedelic drug use and mystical experiences
In the spring of 1953, Huxley had his first experience with the psychedelic drug mescaline. Huxley had initiated a correspondence with Doctor Humphry Osmond, a British psychiatrist then employed in a Canadian institution, and eventually asked him to supply a dose of mescaline; Osmond obliged and supervised Huxley's session in southern California. After the publication of The Doors of Perception, in which he recounted this experience, Huxley and Swami Prabhavananda disagreed about the meaning and importance of the psychedelic drug experience, which may have caused the relationship to cool, but Huxley continued to write articles for the society's journal, lecture at the temple, and attend social functions. Huxley later had an experience on mescaline that he considered more profound than those detailed in The Doors of Perception.
Huxley wrote that "The mystical experience is doubly valuable; it is valuable because it gives the experiencer a better understanding of himself and the world and because it may help him to lead a less self-centered and more creative life."
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628_7 | Eyesight
Differing accounts exist about the details of the quality of Huxley's eyesight at specific points in his life. Circa 1939, Huxley encountered the Bates method, in which he was instructed by Margaret Darst Corbett. In 1940, Huxley relocated from Hollywood to a ranchito in the high desert hamlet of Llano, California, in northern Los Angeles County. Huxley then said that his sight improved dramatically with the Bates Method and the extreme and pure natural lighting of the southwestern American desert. He reported that, for the first time in more than 25 years, he was able to read without glasses and without strain. He even tried driving a car along the dirt road beside the ranch. He wrote a book about his experiences with the Bates Method, The Art of Seeing, which was published in 1942 (U.S.), 1943 (UK). The book contained some generally disputed theories, and its publication created a growing degree of popular controversy about Huxley's eyesight.
It was, and is, widely believed that Huxley was nearly blind since the illness in his teens, despite the partial recovery that had enabled him to study at Oxford. For example, some ten years after publication of The Art of Seeing, in 1952, Bennett Cerf was present when Huxley spoke at a Hollywood banquet, wearing no glasses and apparently reading his paper from the lectern without difficulty: "Then suddenly he faltered—and the disturbing truth became obvious. He wasn't reading his address at all. He had learned it by heart. To refresh his memory he brought the paper closer and closer to his eyes. When it was only an inch or so away he still couldn't read it, and had to fish for a magnifying glass in his pocket to make the typing visible to him. It was an agonising moment."
Brazilian author João Ubaldo Ribeiro, who as a young journalist spent several evenings in the Huxleys' company in the late 1950s, wrote that Huxley had said to him, with a wry smile: "I can hardly see at all. And I don't give a damn, really." | [
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628_8 | On the other hand, Huxley's second wife, Laura Archera, later emphasised in her biographical account, This Timeless Moment: "One of the great achievements of his life: that of having regained his sight." After revealing a letter she wrote to the Los Angeles Times disclaiming the label of Huxley as a "poor fellow who can hardly see" by Walter C. Alvarez, she tempered her statement: "Although I feel it was an injustice to treat Aldous as though he were blind, it is true there were many indications of his impaired vision. For instance, although Aldous did not wear glasses, he would quite often use a magnifying lens." Laura Huxley proceeded to elaborate a few nuances of inconsistency peculiar to Huxley's vision. Her account, in this respect, agrees with the following sample of Huxley's own words from The Art of Seeing: "The most characteristic fact about the functioning of the total organism, or any part of the organism, is that it is not constant, but highly variable." Nevertheless, the topic of Huxley's eyesight has continued to endure similar, significant controversy.
American popular science author Steven Johnson, in his book Mind Wide Open, quotes Huxley about his difficulties with visual encoding: "I am and, for as long as I can remember, I have always been a poor visualizer. Words, even the pregnant words of poets, do not evoke pictures in my mind. No hypnagogic visions greet me on the verge of sleep. When I recall something, the memory does not present itself to me as a vividly seen event or object. By an effort of the will, I can evoke a not very vivid image of what happened yesterday afternoon ..."
Personal life
Huxley married on 10 July 1919 Maria Nys (10 September 1899 – 12 February 1955), a Belgian epidemiologist from Bellem, a village near Aalter, he met at Garsington, Oxfordshire, in 1919. They had one child, Matthew Huxley (19 April 1920 – 10 February 2005), who had a career as an author, anthropologist, and prominent epidemiologist. In 1955, Maria Huxley died of cancer. | [
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628_9 | In 1956, Huxley married Laura Archera (1911–2007), also an author, as well as a violinist and psychotherapist. She wrote This Timeless Moment, a biography of Huxley. She told the story of their marriage through Mary Ann Braubach's 2010 documentary, Huxley on Huxley.
Huxley was diagnosed with laryngeal cancer in 1960; in the years that followed, with his health deteriorating, he wrote the Utopian novel Island, and gave lectures on "Human Potentialities" both at the UCSF Medical Center and at the Esalen Institute. These lectures were fundamental to the beginning of the Human Potential Movement.
Huxley was a close friend of Jiddu Krishnamurti and Rosalind Rajagopal and was involved in the creation of the Happy Valley School, now Besant Hill School of Happy Valley, in Ojai, California.
The most substantial collection of Huxley's few remaining papers, following the destruction of most in the 1961 Bel Air Fire, is at the Library of the University of California, Los Angeles. Some are also at the Stanford University Libraries.
On 9 April 1962 Huxley was informed he was elected Companion of Literature by the Royal Society of Literature, the senior literary organisation in Britain, and he accepted the title via letter on 28 April 1962. The correspondence between Huxley and the society is kept at the Cambridge University Library. The society invited Huxley to appear at a banquet and give a lecture at Somerset House, London, in June 1963. Huxley wrote a draft of the speech he intended to give at the society; however, his deteriorating health meant he was not able to attend.
Death
On his deathbed, unable to speak owing to advanced laryngeal cancer, Huxley made a written request to his wife Laura for "LSD, 100 µg, intramuscular." According to her account of his death in This Timeless Moment, she obliged with an injection at 11:20 a.m. and a second dose an hour later; Huxley died aged 69, at 5:20 p.m. (Los Angeles time), on 22 November 1963. | [
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] |
628_10 | Media coverage of Huxley's death, along with that of fellow British author C. S. Lewis, was overshadowed by the assassination of American President John F. Kennedy on the same day, less than seven hours before Huxley's death. In a 2009 article for New York magazine titled "The Eclipsed Celebrity Death Club", Christopher Bonanos wrote:
This coincidence served as the basis for Peter Kreeft's book Between Heaven and Hell: A Dialog Somewhere Beyond Death with John F. Kennedy, C. S. Lewis, & Aldous Huxley, which imagines a conversation among the three men taking place in Purgatory following their deaths.
Huxley's memorial service took place in London in December 1963; it was led by his elder brother Julian. On 27 October 1971, his ashes were interred in the family grave at the Watts Cemetery, home of the Watts Mortuary Chapel in Compton, Guildford, Surrey, England.
Huxley had been a long-time friend of Russian composer Igor Stravinsky, who dedicated his last orchestral composition to Huxley. What became Variations: Aldous Huxley in memoriam was begun in July 1963, completed in October 1964, and premiered by the Chicago Symphony Orchestra on 17 April 1965.
Awards
1939: James Tait Black Memorial Prize
1959: American Academy of Arts and Letters Award of Merit .
1962: Companion of Literature
Film adaptations of Huxley's work
1950: Prelude to Fame based upon Young Archimedes
1968: Point Counter Point
1971: The Devils
1980: Brave New World
1998: Brave New World
2020: Brave New World
Bibliography
See also
List of peace activists
References
Sources
. Reprinted in Perspectives on Schoenberg and Stravinsky, revised edition, edited by Benjamin Boretz and Edward T. Cone. New York: W. W. Norton, 1972.
Further reading | [
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628_11 | Further reading
Anderson, Jack. 4 July 1982. "Ballet: Suzanne Farrell in Variations Premiere". The New York Times.
Atkins, John. Aldous Huxley: A Literary Study, J. Calder, 1956
Barnes, Clive. 1 April 1966. "Ballet: Still Another Balanchine-Stravinsky Pearl; City Troupe Performs in Premiere Here Variations for Huxley at State Theater". The New York Times, p. 28.
Firchow, Peter. Aldous Huxley: Satirist and Novelist, U of Minnesota P, 1972
Firchow, Peter. The End of Utopia: A Study of Aldous Huxley's Brave New World, Bucknell UP, 1984
Huxley, Aldous. The Human Situation: Aldous Huxley Lectures at Santa Barbara 1959, Flamingo Modern Classic, 1994,
Huxley, Laura Archera. This Timeless Moment, Celestial Arts, 2001,
Meckier, Jerome. Aldous Huxley: Modern Satirical Novelist of Ideas, Firchow and Nugel editors, LIT Verlag Berlin-Hamburg-Münster, 2006,
Morgan, W. John, 'Pacifism or Bourgeois Pacifism? Huxley, Orwell, and Caudwell', Chapter 5 in Morgan, W. John and Guilherme, Alexandre (Eds.),Peace and War-Historical, Philosophical, and Anthropological Perspectives, Palgrave Macmillan, 2020, pp, 71–96. .
Murray, Nicholas. Aldous Huxley, Macmillan, 2003,
Poller, Jake. Aldous Huxley, Reaktion Critical Lives, 2021. .
Poller, Jake. Aldous Huxley and Alternative Spirituality, Brill, 2019. .
Rolo, Charles J. (ed.). The World of Aldous Huxley, Grosset Universal Library, 1947.
Shaw, Jeffrey M. Illusions of Freedom: Thomas Merton and Jacques Ellul on Technology and the Human Condition. Eugene, Oregon: Wipf and Stock. 2014. .
Shadurski, Maxim. The Nationality of Utopia: H. G. Wells, England, and the World State. New York and London: Routledge, 2020. (Chapter 5)
Watt, Conrad (ed.). Aldous Huxley, Routledge, 1997,
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] |
628_12 | External links
Aldous Huxley full interview 1958: The Problems of Survival and Freedom in America
Portraits at the National Portrait Gallery
Raymond Fraser, George Wickes (Spring 1960). "Interview: Aldous Huxley: The Art of Fiction No. 24". The Paris Review.
BBC discussion programme In our time: "Brave New World". Huxley and the novel. 9 April 2009. (Audio, 45 minutes)
BBC In their own words series. 12 October 1958 (video, 12 mins)
"The Ultimate Revolution" (talk at UC Berkeley, 20 March 1962)
Huxley interviewed on The Mike Wallace Interview 18 May 1958 (video)
Centre for Huxley Research
Aldous Huxley Papers at University of California, Los Angeles Library Special Collections
Online editions
1894 births
1963 deaths
20th-century English novelists
20th-century essayists
Alumni of Balliol College, Oxford
Anti-consumerists
Bates method
English emigrants to the United States
20th-century British short story writers
Burials in Surrey
Consciousness researchers and theorists
Deaths from cancer in California
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Duke University faculty
English agnostics
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English expatriates in the United States
English male novelists
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English people of Cornish descent
English satirists
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Futurologists
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Aldous
James Tait Black Memorial Prize recipients
Male essayists
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Perennial philosophy
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Philosophers of literature
Philosophers of mind
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Writers from Los Angeles
Writers from Taos, New Mexico
20th-century English philosophers
Lost Generation writers | [
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630_0 | Ada may refer to:
Places
Africa
Ada Foah or Ada, Ghana, a town
Ada (Ghana parliament constituency)
Ada, Osun, a town in Osun State, Nigeria
Asia
Adeh, Urmia, also known as Ada, a village in West Azerbaijan Province
Ada, Karaman, a village in Karaman Province, Turkey
Australia and New Zealand
Ada River (disambiguation), three rivers
Europe
Ada, Bosnia and Herzegovina, a village
Ada, Croatia, a village
Ada, Serbia, a town and municipality
Ada Ciganlija or Ada, a river island artificially turned into a peninsula in Belgrade, Serbia
North America
United States
Ada, Alabama, an unincorporated community
Ada County, Idaho
Ada, Kansas, an unincorporated community
Ada Township, Michigan
Ada, Minnesota, a city
Ada Township, Dickey County, North Dakota
Ada, Ohio, a village
Ada, Oklahoma, a city
Ada, Oregon, an unincorporated community
Ada Township, Perkins County, South Dakota
Ada, West Virginia, an unincorporated community
Ada, Wisconsin, an unincorporated community
Outer space
523 Ada, an asteroid
Film and television
Ada TV, a television channel in the Turkish Republic of Northern Cyprus
Ada (1961 film), a 1961 film by Daniel Mann
Ada (2019 film), a short biopic about Ada Lovelace
Ada... A Way of Life, a 2008 Bollywood musical by Tanvir Ahmed
Ada (dog actor), a dog that played Colin on the sitcom Spaced
Ada, one of the main characters in 1991 movie Armour of God II: Operation Condor
Biology
Ada (plant), a genus of orchids
Adenosine deaminase, an enzyme involved in purine metabolism
Ada (protein), an enzyme induced by treatment of bacterial cells
Computer science
Ada (programming language), programming language based on Pascal
Ada (computer virus)
Air travel
Ada Air, a regional airline based in Tirana, Albania
Ada International Airport or Saipan International Airport, Saipan Island, Northern Mariana Islands
Aerolínea de Antioquia, a Colombian airline
Airline Deregulation Act, a 1978 US bill removing governmental control from commercial aviation | [
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630_1 | Schools
Ada, the National College for Digital Skills, a further education college in Tottenham Hale, London
Ada High School (Ohio), Ada, Ohio
Ada High School (Oklahoma), Ada, Oklahoma
People
Ada (name), a feminine given name and a surname, including a list of people and fictional characters
Ada Lovelace (1815–1852), computer scientist sometimes regarded as the first computer programmer
Other uses
List of tropical storms named Ada
Ada (food), a traditional Kerala delicacy
Ada, the cryptocurrency of the Cardano blockchain platform
Ada Bridge, Belgrade, Serbia
, a cargo vessel built for the London and South Western Railway
Ada (ship), a wooden ketch, wrecked near Newcastle, New South Wales, Australia
Ada or Ardor: A Family Chronicle, novel by Vladimir Nabokov
Dangme language, spoken in Ghana (ISO 639-2 and 639-3 code "ada")
Ada Health GmbH, a symptom checker app
See also
ADA (disambiguation)
Ada regulon, an Escherichia coli adaptive response protein
Adah (disambiguation)
Adha (disambiguation)
Ada'a, a woreda in the Oromia Region of Ethiopia
Ade (disambiguation)
USS Little Ada (1864), a steamer captured by the Union Navy during the American Civil War | [
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632_0 | Aberdeen is a city in Scotland, United Kingdom.
Aberdeen may also refer to:
Places
Africa
Aberdeen, Sierra Leone
Aberdeen, Eastern Cape, South Africa
Asia
Hong Kong
Aberdeen, Hong Kong, an area and town on southwest Hong Kong Island
Aberdeen Channel, a channel between Ap Lei Chau (Aberdeen Island) and Nam Long Shan on the Hong Kong Island in Hong Kong
Aberdeen Country Park, a country park in Hong Kong Island
Aberdeen floating village, at Aberdeen Harbour, containing approximately 600 junks, which house an estimated 6,000 people
Aberdeen Harbour, a harbour between Aberdeen, Hong Kong and Ap Lei Chau (Aberdeen Island)
Aberdeen Tunnel, a tunnel in Hong Kong Island
Aberdeen Tunnel Underground Laboratory, an underground particle physics laboratory in Hong Kong Island
Ap Lei Chau or Aberdeen Island, an island of Hong Kong
Aberdeen (constituency), a constituency of Southern District Council
India
Aberdeen Bazaar, a shopping centre in Port Blair, South Andaman Island
Sri Lanka
Aberdeen Falls, a waterfall in Sri Lanka
Australia
Aberdeen, New South Wales
Aberdeen, South Australia, one of the early townships that merged in 1940 to create the town of Burra
Aberdeen, Tasmania, a suburb of the City of Devonport
Caribbean
Aberdeen, Jamaica, a town in Saint Elizabeth, Jamaica
Europe
Aberdeen (Parliament of Scotland constituency)
Aberdeen (UK Parliament constituency) 1832-1885
Aberdeen Burghs (UK Parliament constituency) 1801-1832
Aberdeen Central (Scottish Parliament constituency)
Aberdeen Central (UK Parliament constituency)
Aberdeen Donside (Scottish Parliament constituency)
County of Aberdeen, a historic county of Scotland whose county town was Aberdeen
Old Aberdeen, a part of the city of Aberdeen in Scotland
North America | [
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632_1 | North America
Canada
Aberdeen, community in the township of Champlain, Prescott and Russell County, Ontario
Aberdeen, Abbotsford, a neighbourhood in the City of Abbotsford, British Columbia
Aberdeen Centre, a shopping mall in Richmond, British Columbia
Aberdeen, Grey County, Ontario
Aberdeen, Kamloops, an area in the City of Kamloops, British Columbia
Aberdeen Lake (Nunavut), a lake in Kivalliq Region, Nunavut, Canada
Aberdeen, Nova Scotia, part of the Municipality of Inverness County, Nova Scotia
Aberdeen Parish, New Brunswick
Rural Municipality of Aberdeen No. 373, Saskatchewan
Aberdeen, Saskatchewan
Aberdeen Bay, a bay between southern Baffin Island and north-eastern Hector Island in the Nunavut territory
Aberdeen Township, Quebec, until 1960 part of Sheen-Esher-Aberdeen-et-Malakoff, now part of Rapides-des-Joachims, Quebec
Aberdeen River, a tributary of rivière aux Castors Noirs in Mauricie, Québec
New Aberdeen, Nova Scotia
United States
Aberdeen, Arkansas
Aberdeen, Florida
Aberdeen, Georgia
Aberdeen, Idaho
Aberdeen, Ohio County, Indiana
Aberdeen, Porter County, Indiana
Aberdeen, Kentucky
Aberdeen, Maryland
Aberdeen Proving Ground, a United States Army facility located near Aberdeen, Maryland
Aberdeen, Massachusetts, a neighborhood of Brighton, Boston
Aberdeen, Mississippi
Aberdeen Lake (Mississippi), a lake in northeast Mississippi on the Tennessee-Tombigbee Waterway, close to Aberdeen, Mississippi
Aberdeen Township, New Jersey
Aberdeen, North Carolina
Aberdeen Historic District (Aberdeen, North Carolina)
Aberdeen, Ohio
Aberdeen, South Dakota
Aberdeen Historic District (Aberdeen, South Dakota)
Aberdeen, Texas
Aberdeen (Disputanta, Virginia)
Aberdeen Gardens (Hampton, Virginia)
Aberdeen, Washington
Aberdeen Gardens, Washington
Aberdeen, West Virginia
Business
Abrdn, formerly Standard Life Aberdeen
Aberdeen Asset Management
Education
Aberdeen Business School
Aberdeen College, formerly one of the largest further education colleges in Scotland, merged with Banff & Buchan College to form North East Scotland College
Aberdeen Grammar School, Aberdeen, Scotland
Aberdeen Hall, a university-preparatory school in Kelowna, British Columbia, Canada
Aberdeen High School (disambiguation)
King's College, Aberdeen
University of Aberdeen, a public research university in the city of Aberdeen | [
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] |
632_2 | Entertainment
Aberdeen (2000 film), a 2000 Norwegian-British film directed by Hans Petter Moland, starring Stellan Skarsgård and Lena Headey
Aberdeen (2014 film), a 2014 Hong Kong film starring Louis Koo
Aberdeen (band), an American rock band
Aberdeen (song), a song by Cage The Elephant
Aberdeen City (band), Boston based indie/alternative rock band
Other transportation
Aberdeen Airport (disambiguation)
Aberdeen Lock and Dam, one of four lock and dam structures on the Tennessee-Tombigbee Waterway
Rail
Aberdeen, Carolina and Western Railway, a short-line railroad operating in North Carolina
Aberdeen and Rockfish Railroad, a short-line railroad operating in North Carolina
Aberdeen Corporation Tramways
Aberdeen Line (disambiguation)
Aberdeen station (disambiguation)
Dundee and Perth and Aberdeen Junction Railway, a later name of the Dundee and Perth Railway
Shipping
Aberdeen Line, a British shipping company founded in 1825
, one of several ships by that name
, a sloop of the British Royal Navy that served between 1936 and 1948
, a merchant ship operated during the latter stages of World War II, later commissioned as the USS Altair
Sports
Aberdeen Dad Vail Regatta, the largest regular intercollegiate rowing event in the United States, named after its sponsor, Aberdeen Asset Management
Aberdeen F.C. (disambiguation)
Aberdeen GSFP RFC, an amateur rugby union club based in Aberdeen
Aberdeen IronBirds, a minor league baseball team affiliated with the Baltimore Orioles
Aberdeen L.F.C., a women's football team affiliated with Aberdeen F.C.
See also
Aberdeen Act
Aberdeen Angus, a Scottish breed of small beef cattle
Aberdeen Central (disambiguation)
Aberdeen Gardens (disambiguation)
Aberdeen Historic District (disambiguation)
Aberdeen Hospital (disambiguation)
Aberdeen Quarry, a granite quarry in Colorado
Battle of Aberdeen (disambiguation)
Diocese of Aberdeen and Orkney, one of the seven dioceses of the Scottish Episcopal Church
Etymology of Aberdeen
Marquess of Aberdeen and Temair, a title in the Peerage of the United Kingdom | [
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633_0 | Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular microalgae, such as Chlorella, Prototheca and the diatoms, to multicellular forms, such as the giant kelp, a large brown alga which may grow up to in length. Most are aquatic and autotrophic (they generate food internally) and lack many of the distinct cell and tissue types, such as stomata, xylem and phloem that are found in land plants. The largest and most complex marine algae are called seaweeds, while the most complex freshwater forms are the Charophyta, a division of green algae which includes, for example, Spirogyra and stoneworts.
No definition of algae is generally accepted. One definition is that algae "have chlorophyll as their primary photosynthetic pigment and lack a sterile covering of cells around their reproductive cells". Likewise, the colorless Prototheca under Chlorophyta are all devoid of any chlorophyll. Although cyanobacteria are often referred to as "blue-green algae", most authorities exclude all prokaryotes from the definition of algae.
Algae constitute a polyphyletic group since they do not include a common ancestor, and although their plastids seem to have a single origin, from cyanobacteria, they were acquired in different ways. Green algae are examples of algae that have primary chloroplasts derived from endosymbiotic cyanobacteria. Diatoms and brown algae are examples of algae with secondary chloroplasts derived from an endosymbiotic red alga. Algae exhibit a wide range of reproductive strategies, from simple asexual cell division to complex forms of sexual reproduction. | [
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633_1 | Algae lack the various structures that characterize land plants, such as the phyllids (leaf-like structures) of bryophytes, rhizoids of nonvascular plants, and the roots, leaves, and other organs found in tracheophytes (vascular plants). Most are phototrophic, although some are mixotrophic, deriving energy both from photosynthesis and uptake of organic carbon either by osmotrophy, myzotrophy, or phagotrophy. Some unicellular species of green algae, many golden algae, euglenids, dinoflagellates, and other algae have become heterotrophs (also called colorless or apochlorotic algae), sometimes parasitic, relying entirely on external energy sources and have limited or no photosynthetic apparatus. Some other heterotrophic organisms, such as the apicomplexans, are also derived from cells whose ancestors possessed plastids, but are not traditionally considered as algae. Algae have photosynthetic machinery ultimately derived from cyanobacteria that produce oxygen as a by-product of photosynthesis, unlike other photosynthetic bacteria such as purple and green sulfur bacteria. Fossilized filamentous algae from the Vindhya basin have been dated back to 1.6 to 1.7 billion years ago.
Because of the wide range of types of algae, they have increasing different industrial and traditional applications in human society. Traditional seaweed farming practices have existed for thousands of years and have strong traditions in East Asia food cultures. More modern algaculture applications extend the food traditions for other applications include cattle feed, using algae for bioremediation or pollution control, transforming sunlight into algae fuels or other chemicals used in industrial processes, and in medical and scientific applications. A 2020 review, found that these applications of algae could play an important role in carbon sequestration in order to mitigate climate change while providing valuable value-add products for global economies.
Etymology and study
The singular is the Latin word for 'seaweed' and retains that meaning in English. The etymology is obscure. Although some speculate that it is related to Latin , 'be cold', no reason is known to associate seaweed with temperature. A more likely source is , 'binding, entwining'. | [
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633_2 | The Ancient Greek word for 'seaweed' was (), which could mean either the seaweed (probably red algae) or a red dye derived from it. The Latinization, , meant primarily the cosmetic rouge. The etymology is uncertain, but a strong candidate has long been some word related to the Biblical (), 'paint' (if not that word itself), a cosmetic eye-shadow used by the ancient Egyptians and other inhabitants of the eastern Mediterranean. It could be any color: black, red, green, or blue.
Accordingly, the modern study of marine and freshwater algae is called either phycology or algology, depending on whether the Greek or Latin root is used. The name fucus appears in a number of taxa.
Classifications
The committee on the International Code of Botanical Nomenclature has recommended certain suffixes for use in the classification of algae. These are -phyta for division, -phyceae for class, -phycideae for subclass, -ales for order, -inales for suborder, -aceae for family, -oidease for subfamily, a Greek-based name for genus, and a Latin-based name for species.
Algal characteristics basic to primary classification
The primary classification of algae is based on certain morphological features. The chief among these are (a) pigment constitution of the cell, (b) chemical nature of stored food materials, (c) kind, number, point of insertion and relative length of the flagella on the motile cell, (d) chemical composition of cell wall and (e) presence or absence of a definitely organized nucleus in the cell or any other significant details of cell structure.
History of classification of algae
Although Carolus Linnaeus (1754) included algae along with lichens in his 25th class Cryptogamia, he did not elaborate further on the classification of algae. | [
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633_3 | Jean Pierre Étienne Vaucher (1803) was perhaps the first to propose a system of classification of algae, and he recognized three groups, Conferves, Ulves, and Tremelles. While Johann Heinrich Friedrich Link (1820) classified algae on the basis of the colour of the pigment and structure, William Henry Harvey (1836) proposed a system of classification on the basis of the habitat and the pigment. J. G. Agardh (1849–1898) divided algae into six orders: Diatomaceae, Nostochineae, Confervoideae, Ulvaceae, Floriadeae and Fucoideae. Around 1880, algae along with fungi were grouped under Thallophyta, a division created by Eichler (1836). Encouraged by this, Adolf Engler and Karl A. E. Prantl (1912) proposed a revised scheme of classification of algae and included fungi in algae as they were of opinion that fungi have been derived from algae. The scheme proposed by Engler and Prantl is summarised as follows:
Schizophyta
Phytosarcodina
Flagellata
Dinoflagellata
Bacillariophyta
Conjugatae
Chlorophyceae
Charophyta
Phaeophyceae
Rhodophyceae
Eumycetes (Fungi)
The algae contain chloroplasts that are similar in structure to cyanobacteria. Chloroplasts contain circular DNA like that in cyanobacteria and are interpreted as representing reduced endosymbiotic cyanobacteria. However, the exact origin of the chloroplasts is different among separate lineages of algae, reflecting their acquisition during different endosymbiotic events. The table below describes the composition of the three major groups of algae. Their lineage relationships are shown in the figure in the upper right. Many of these groups contain some members that are no longer photosynthetic. Some retain plastids, but not chloroplasts, while others have lost plastids entirely.
Phylogeny based on plastid not nucleocytoplasmic genealogy: | [
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633_4 | Linnaeus, in Species Plantarum (1753), the starting point for modern botanical nomenclature, recognized 14 genera of algae, of which only four are currently considered among algae. In Systema Naturae, Linnaeus described the genera Volvox and Corallina, and a species of Acetabularia (as Madrepora), among the animals.
In 1768, Samuel Gottlieb Gmelin (1744–1774) published the Historia Fucorum, the first work dedicated to marine algae and the first book on marine biology to use the then new binomial nomenclature of Linnaeus. It included elaborate illustrations of seaweed and marine algae on folded leaves.
W. H. Harvey (1811–1866) and Lamouroux (1813) were the first to divide macroscopic algae into four divisions based on their pigmentation. This is the first use of a biochemical criterion in plant systematics. Harvey's four divisions are: red algae (Rhodospermae), brown algae (Melanospermae), green algae (Chlorospermae), and Diatomaceae.
At this time, microscopic algae were discovered and reported by a different group of workers (e.g., O. F. Müller and Ehrenberg) studying the Infusoria (microscopic organisms). Unlike macroalgae, which were clearly viewed as plants, microalgae were frequently considered animals because they are often motile. Even the nonmotile (coccoid) microalgae were sometimes merely seen as stages of the lifecycle of plants, macroalgae, or animals.
Although used as a taxonomic category in some pre-Darwinian classifications, e.g., Linnaeus (1753), de Jussieu (1789), Horaninow (1843), Agassiz (1859), Wilson & Cassin (1864), in further classifications, the "algae" are seen as an artificial, polyphyletic group. | [
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633_5 | Throughout the 20th century, most classifications treated the following groups as divisions or classes of algae: cyanophytes, rhodophytes, chrysophytes, xanthophytes, bacillariophytes, phaeophytes, pyrrhophytes (cryptophytes and dinophytes), euglenophytes, and chlorophytes. Later, many new groups were discovered (e.g., Bolidophyceae), and others were splintered from older groups: charophytes and glaucophytes (from chlorophytes), many heterokontophytes (e.g., synurophytes from chrysophytes, or eustigmatophytes from xanthophytes), haptophytes (from chrysophytes), and chlorarachniophytes (from xanthophytes).
With the abandonment of plant-animal dichotomous classification, most groups of algae (sometimes all) were included in Protista, later also abandoned in favour of Eukaryota. However, as a legacy of the older plant life scheme, some groups that were also treated as protozoans in the past still have duplicated classifications (see ambiregnal protists). | [
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633_6 | Some parasitic algae (e.g., the green algae Prototheca and Helicosporidium, parasites of metazoans, or Cephaleuros, parasites of plants) were originally classified as fungi, sporozoans, or protistans of incertae sedis, while others (e.g., the green algae Phyllosiphon and Rhodochytrium, parasites of plants, or the red algae Pterocladiophila and Gelidiocolax mammillatus, parasites of other red algae, or the dinoflagellates Oodinium, parasites of fish) had their relationship with algae conjectured early. In other cases, some groups were originally characterized as parasitic algae (e.g., Chlorochytrium), but later were seen as endophytic algae. Some filamentous bacteria (e.g., Beggiatoa) were originally seen as algae. Furthermore, groups like the apicomplexans are also parasites derived from ancestors that possessed plastids, but are not included in any group traditionally seen as algae.
Relationship to land plants
The first land plants probably evolved from shallow freshwater charophyte algae much like Chara almost 500 million years ago. These probably had an isomorphic alternation of generations and were probably filamentous. Fossils of isolated land plant spores suggest land plants may have been around as long as 475 million years ago.
Morphology
A range of algal morphologies is exhibited, and convergence of features in unrelated groups is common. The only groups to exhibit three-dimensional multicellular thalli are the reds and browns, and some chlorophytes. Apical growth is constrained to subsets of these groups: the florideophyte reds, various browns, and the charophytes. The form of charophytes is quite different from those of reds and browns, because they have distinct nodes, separated by internode 'stems'; whorls of branches reminiscent of the horsetails occur at the nodes. Conceptacles are another polyphyletic trait; they appear in the coralline algae and the Hildenbrandiales, as well as the browns. | [
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633_7 | Most of the simpler algae are unicellular flagellates or amoeboids, but colonial and nonmotile forms have developed independently among several of the groups. Some of the more common organizational levels, more than one of which may occur in the lifecycle of a species, are
Colonial: small, regular groups of motile cells
Capsoid: individual non-motile cells embedded in mucilage
Coccoid: individual non-motile cells with cell walls
Palmelloid: nonmotile cells embedded in mucilage
Filamentous: a string of nonmotile cells connected together, sometimes branching
Parenchymatous: cells forming a thallus with partial differentiation of tissues
In three lines, even higher levels of organization have been reached, with full tissue differentiation. These are the brown algae,—some of which may reach 50 m in length (kelps)—the red algae, and the green algae. The most complex forms are found among the charophyte algae (see Charales and Charophyta), in a lineage that eventually led to the higher land plants. The innovation that defines these nonalgal plants is the presence of female reproductive organs with protective cell layers that protect the zygote and developing embryo. Hence, the land plants are referred to as the Embryophytes. | [
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633_8 | Turfs
The term algal turf is commonly used but poorly defined. Algal turfs are thick, carpet-like beds of seaweed that retain sediment and compete with foundation species like corals and kelps, and they are usually less than 15 cm tall. Such a turf may consist of one or more species, and will generally cover an area in the order of a square metre or more. Some common characteristics are listed:
Algae that form aggregations that have been described as turfs include diatoms, cyanobacteria, chlorophytes, phaeophytes and rhodophytes. Turfs are often composed of numerous species at a wide range of spatial scales, but monospecific turfs are frequently reported.
Turfs can be morphologically highly variable over geographic scales and even within species on local scales and can be difficult to identify in terms of the constituent species.
Turfs have been defined as short algae, but this has been used to describe height ranges from less than 0.5 cm to more than 10 cm. In some regions, the descriptions approached heights which might be described as canopies (20 to 30 cm).
Physiology
Many algae, particularly members of the Characeae species, have served as model experimental organisms to understand the mechanisms of the water permeability of membranes, osmoregulation, turgor regulation, salt tolerance, cytoplasmic streaming, and the generation of action potentials.
Phytohormones are found not only in higher plants, but in algae, too.
Symbiotic algae
Some species of algae form symbiotic relationships with other organisms. In these symbioses, the algae supply photosynthates (organic substances) to the host organism providing protection to the algal cells. The host organism derives some or all of its energy requirements from the algae. Examples are:
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633_9 | Lichens
Lichens are defined by the International Association for Lichenology to be "an association of a fungus and a photosynthetic symbiont resulting in a stable vegetative body having a specific structure". The fungi, or mycobionts, are mainly from the Ascomycota with a few from the Basidiomycota. In nature they do not occur separate from lichens. It is unknown when they began to associate. One mycobiont associates with the same phycobiont species, rarely two, from the green algae, except that alternatively, the mycobiont may associate with a species of cyanobacteria (hence "photobiont" is the more accurate term). A photobiont may be associated with many different mycobionts or may live independently; accordingly, lichens are named and classified as fungal species. The association is termed a morphogenesis because the lichen has a form and capabilities not possessed by the symbiont species alone (they can be experimentally isolated). The photobiont possibly triggers otherwise latent genes in the mycobiont.
Trentepohlia is an example of a common green alga genus worldwide that can grow on its own or be lichenised. Lichen thus share some of the habitat and often similar appearance with specialized species of algae (aerophytes) growing on exposed surfaces such as tree trunks and rocks and sometimes discoloring them.
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633_10 | Coral reefs
Coral reefs are accumulated from the calcareous exoskeletons of marine invertebrates of the order Scleractinia (stony corals). These animals metabolize sugar and oxygen to obtain energy for their cell-building processes, including secretion of the exoskeleton, with water and carbon dioxide as byproducts. Dinoflagellates (algal protists) are often endosymbionts in the cells of the coral-forming marine invertebrates, where they accelerate host-cell metabolism by generating sugar and oxygen immediately available through photosynthesis using incident light and the carbon dioxide produced by the host. Reef-building stony corals (hermatypic corals) require endosymbiotic algae from the genus Symbiodinium to be in a healthy condition. The loss of Symbiodinium from the host is known as coral bleaching, a condition which leads to the deterioration of a reef.
Sea sponges
Endosymbiontic green algae live close to the surface of some sponges, for example, breadcrumb sponges (Halichondria panicea). The alga is thus protected from predators; the sponge is provided with oxygen and sugars which can account for 50 to 80% of sponge growth in some species. | [
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633_11 | Lifecycle
Rhodophyta, Chlorophyta, and Heterokontophyta, the three main algal divisions, have lifecycles which show considerable variation and complexity. In general, an asexual phase exists where the seaweed's cells are diploid, a sexual phase where the cells are haploid, followed by fusion of the male and female gametes. Asexual reproduction permits efficient population increases, but less variation is possible. Commonly, in sexual reproduction of unicellular and colonial algae, two specialized, sexually compatible, haploid gametes make physical contact and fuse to form a zygote. To ensure a successful mating, the development and release of gametes is highly synchronized and regulated; pheromones may play a key role in these processes. Sexual reproduction allows for more variation and provides the benefit of efficient recombinational repair of DNA damages during meiosis, a key stage of the sexual cycle. However, sexual reproduction is more costly than asexual reproduction. Meiosis has been shown to occur in many different species of algae.
Numbers
The Algal Collection of the US National Herbarium (located in the National Museum of Natural History) consists of approximately 320,500 dried specimens, which, although not exhaustive (no exhaustive collection exists), gives an idea of the order of magnitude of the number of algal species (that number remains unknown). Estimates vary widely. For example, according to one standard textbook, in the British Isles the UK Biodiversity Steering Group Report estimated there to be 20,000 algal species in the UK. Another checklist reports only about 5,000 species. Regarding the difference of about 15,000 species, the text concludes: "It will require many detailed field surveys before it is possible to provide a reliable estimate of the total number of species ..." | [
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633_12 | Regional and group estimates have been made, as well:
5,000–5,500 species of red algae worldwide
"some 1,300 in Australian Seas"
400 seaweed species for the western coastline of South Africa, and 212 species from the coast of KwaZulu-Natal. Some of these are duplicates, as the range extends across both coasts, and the total recorded is probably about 500 species. Most of these are listed in List of seaweeds of South Africa. These exclude phytoplankton and crustose corallines.
669 marine species from California (US)
642 in the check-list of Britain and Ireland
and so on, but lacking any scientific basis or reliable sources, these numbers have no more credibility than the British ones mentioned above. Most estimates also omit microscopic algae, such as phytoplankton.
The most recent estimate suggests 72,500 algal species worldwide.
Distribution
The distribution of algal species has been fairly well studied since the founding of phytogeography in the mid-19th century. Algae spread mainly by the dispersal of spores analogously to the dispersal of Plantae by seeds and spores. This dispersal can be accomplished by air, water, or other organisms. Due to this, spores can be found in a variety of environments: fresh and marine waters, air, soil, and in or on other organisms. Whether a spore is to grow into an organism depends on the combination of the species and the environmental conditions where the spore lands.
The spores of freshwater algae are dispersed mainly by running water and wind, as well as by living carriers. However, not all bodies of water can carry all species of algae, as the chemical composition of certain water bodies limits the algae that can survive within them. Marine spores are often spread by ocean currents. Ocean water presents many vastly different habitats based on temperature and nutrient availability, resulting in phytogeographic zones, regions, and provinces. | [
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633_13 | To some degree, the distribution of algae is subject to floristic discontinuities caused by geographical features, such as Antarctica, long distances of ocean or general land masses. It is, therefore, possible to identify species occurring by locality, such as "Pacific algae" or "North Sea algae". When they occur out of their localities, hypothesizing a transport mechanism is usually possible, such as the hulls of ships. For example, Ulva reticulata and U. fasciata travelled from the mainland to Hawaii in this manner.
Mapping is possible for select species only: "there are many valid examples of confined distribution patterns." For example, Clathromorphum is an arctic genus and is not mapped far south of there. However, scientists regard the overall data as insufficient due to the "difficulties of undertaking such studies."
Ecology
Algae are prominent in bodies of water, common in terrestrial environments, and are found in unusual environments, such as on snow and ice. Seaweeds grow mostly in shallow marine waters, under deep; however, some such as Navicula pennata have been recorded to a depth of . A type of algae, Ancylonema nordenskioeldii, was found in Greenland in areas known as the 'Dark Zone', which caused an increase in the rate of melting ice sheet. Same algae was found in the Italian Alps, after pink ice appeared on parts of the Presena glacier.
The various sorts of algae play significant roles in aquatic ecology. Microscopic forms that live suspended in the water column (phytoplankton) provide the food base for most marine food chains. In very high densities (algal blooms), these algae may discolor the water and outcompete, poison, or asphyxiate other life forms.
Algae can be used as indicator organisms to monitor pollution in various aquatic systems. In many cases, algal metabolism is sensitive to various pollutants. Due to this, the species composition of algal populations may shift in the presence of chemical pollutants. To detect these changes, algae can be sampled from the environment and maintained in laboratories with relative ease. | [
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633_14 | On the basis of their habitat, algae can be categorized as: aquatic (planktonic, benthic, marine, freshwater, lentic, lotic), terrestrial, aerial (subaerial), lithophytic, halophytic (or euryhaline), psammon, thermophilic, cryophilic, epibiont (epiphytic, epizoic), endosymbiont (endophytic, endozoic), parasitic, calcifilic or lichenic (phycobiont).
Cultural associations
In classical Chinese, the word is used both for "algae" and (in the modest tradition of the imperial scholars) for "literary talent". The third island in Kunming Lake beside the Summer Palace in Beijing is known as the Zaojian Tang Dao, which thus simultaneously means "Island of the Algae-Viewing Hall" and "Island of the Hall for Reflecting on Literary Talent".
Cultivation
Seaweed farming
Bioreactors
Uses
Agar
Agar, a gelatinous substance derived from red algae, has a number of commercial uses. It is a good medium on which to grow bacteria and fungi, as most microorganisms cannot digest agar.
Alginates
Alginic acid, or alginate, is extracted from brown algae. Its uses range from gelling agents in food, to medical dressings. Alginic acid also has been used in the field of biotechnology as a biocompatible medium for cell encapsulation and cell immobilization. Molecular cuisine is also a user of the substance for its gelling properties, by which it becomes a delivery vehicle for flavours.
Between 100,000 and 170,000 wet tons of Macrocystis are harvested annually in New Mexico for alginate extraction and abalone feed.
Energy source
To be competitive and independent from fluctuating support from (local) policy on the long run, biofuels should equal or beat the cost level of fossil fuels. Here, algae-based fuels hold great promise, directly related to the potential to produce more biomass per unit area in a year than any other form of biomass. The break-even point for algae-based biofuels is estimated to occur by 2025. | [
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633_15 | Fertilizer
For centuries, seaweed has been used as a fertilizer; George Owen of Henllys writing in the 16th century referring to drift weed in South Wales:
Today, algae are used by humans in many ways; for example, as fertilizers, soil conditioners, and livestock feed. Aquatic and microscopic species are cultured in clear tanks or ponds and are either harvested or used to treat effluents pumped through the ponds. Algaculture on a large scale is an important type of aquaculture in some places. Maerl is commonly used as a soil conditioner.
Nutrition
Naturally growing seaweeds are an important source of food, especially in Asia, leading some to label them as superfoods. They provide many vitamins including: A, B1, B2, B6, niacin, and C, and are rich in iodine, potassium, iron, magnesium, and calcium. In addition, commercially cultivated microalgae, including both algae and cyanobacteria, are marketed as nutritional supplements, such as spirulina, Chlorella and the vitamin-C supplement from Dunaliella, high in beta-carotene.
Algae are national foods of many nations: China consumes more than 70 species, including fat choy, a cyanobacterium considered a vegetable; Japan, over 20 species such as nori and aonori; Ireland, dulse; Chile, cochayuyo. Laver is used to make laver bread in Wales, where it is known as ; in Korea, . It is also used along the west coast of North America from California to British Columbia, in Hawaii and by the Māori of New Zealand. Sea lettuce and badderlocks are salad ingredients in Scotland, Ireland, Greenland, and Iceland. Algae is being considered a potential solution for world hunger problem. | [
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633_16 | Two popular forms of algae are used in cuisine:
Chlorella: This form of alga is found in freshwater and contains photosynthetic pigments in its chloroplast. It is high in iron, zinc, magnesium, vitamin B2 and Omega-3 Fatty acids.
Furthermore, it contains all nine of the essential amino acids the body does not produce on its own
Spirulina: Known otherwise as a cyanobacterium (a prokaryote, incorrectly referred to as a "blue-green alga"), contains 10% more protein than Chlorella as well as more thiamine and copper.
The oils from some algae have high levels of unsaturated fatty acids. For example, Parietochloris incisa is very high in arachidonic acid, where it reaches up to 47% of the triglyceride pool. Some varieties of algae favored by vegetarianism and veganism contain the long-chain, essential omega-3 fatty acids, docosahexaenoic acid (DHA) and eicosapentaenoic acid (EPA). Fish oil contains the omega-3 fatty acids, but the original source is algae (microalgae in particular), which are eaten by marine life such as copepods and are passed up the food chain. Algae have emerged in recent years as a popular source of omega-3 fatty acids for vegetarians who cannot get long-chain EPA and DHA from other vegetarian sources such as flaxseed oil, which only contains the short-chain alpha-linolenic acid (ALA).
Pollution control
Sewage can be treated with algae, reducing the use of large amounts of toxic chemicals that would otherwise be needed.
Algae can be used to capture fertilizers in runoff from farms. When subsequently harvested, the enriched algae can be used as fertilizer.
Aquaria and ponds can be filtered using algae, which absorb nutrients from the water in a device called an algae scrubber, also known as an algae turf scrubber. | [
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633_17 | Agricultural Research Service scientists found that 60–90% of nitrogen runoff and 70–100% of phosphorus runoff can be captured from manure effluents using a horizontal algae scrubber, also called an algal turf scrubber (ATS). Scientists developed the ATS, which consists of shallow, 100-foot raceways of nylon netting where algae colonies can form, and studied its efficacy for three years. They found that algae can readily be used to reduce the nutrient runoff from agricultural fields and increase the quality of water flowing into rivers, streams, and oceans. Researchers collected and dried the nutrient-rich algae from the ATS and studied its potential as an organic fertilizer. They found that cucumber and corn seedlings grew just as well using ATS organic fertilizer as they did with commercial fertilizers. Algae scrubbers, using bubbling upflow or vertical waterfall versions, are now also being used to filter aquaria and ponds.
Polymers
Various polymers can be created from algae, which can be especially useful in the creation of bioplastics. These include hybrid plastics, cellulose-based plastics, poly-lactic acid, and bio-polyethylene. Several companies have begun to produce algae polymers commercially, including for use in flip-flops and in surf boards.
Bioremediation
The alga Stichococcus bacillaris has been seen to colonize silicone resins used at archaeological sites; biodegrading the synthetic substance.
Pigments
The natural pigments (carotenoids and chlorophylls) produced by algae can be used as alternatives to chemical dyes and coloring agents.
The presence of some individual algal pigments, together with specific pigment concentration ratios, are taxon-specific: analysis of their concentrations with various analytical methods, particularly high-performance liquid chromatography, can therefore offer deep insight into the taxonomic composition and relative abundance of natural algae populations in sea water samples.
Stabilizing substances
Carrageenan, from the red alga Chondrus crispus, is used as a stabilizer in milk products.
Additional images
See also
AlgaeBase
AlgaePARC
Eutrophication
Iron fertilization
Marimo algae
Microbiofuels
Microphyte
Photobioreactor
Phycotechnology
Plant
Toxoid – anatoxin | [
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] |
633_18 | References
Bibliography
General
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External links
– a database of all algal names including images, nomenclature, taxonomy, distribution, bibliography, uses, extracts
EnAlgae
Endosymbiotic events
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634_0 | Analysis of variance (ANOVA) is a collection of statistical models and their associated estimation procedures (such as the "variation" among and between groups) used to analyze the differences among means. ANOVA was developed by the statistician Ronald Fisher. ANOVA is based on the law of total variance, where the observed variance in a particular variable is partitioned into components attributable to different sources of variation. In its simplest form, ANOVA provides a statistical test of whether two or more population means are equal, and therefore generalizes the t-test beyond two means.
History
While the analysis of variance reached fruition in the 20th century, antecedents extend centuries into the past according to Stigler. These include hypothesis testing, the partitioning of sums of squares, experimental techniques and the additive model. Laplace was performing hypothesis testing in the 1770s. Around 1800, Laplace and Gauss developed the least-squares method for combining observations, which improved upon methods then used in astronomy and geodesy. It also initiated much study of the contributions to sums of squares. Laplace knew how to estimate a variance from a residual (rather than a total) sum of squares. By 1827, Laplace was using least squares methods to address ANOVA problems regarding measurements of atmospheric tides. Before 1800, astronomers had isolated observational errors resulting
from reaction times (the "personal equation") and had developed methods of reducing the errors. The experimental methods used in the study of the personal equation were later accepted by the emerging field of psychology which developed strong (full factorial) experimental methods to which randomization and blinding were soon added. An eloquent non-mathematical explanation of the additive effects model was available in 1885.
Ronald Fisher introduced the term variance and proposed its formal analysis in a 1918 article The Correlation Between Relatives on the Supposition of Mendelian Inheritance. His first application of the analysis of variance was published in 1921. Analysis of variance became widely known after being included in Fisher's 1925 book Statistical Methods for Research Workers.
Randomization models were developed by several researchers. The first was published in Polish by Jerzy Neyman in 1923. | [
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634_1 | Example
The analysis of variance can be used to describe otherwise complex relations among variables. A dog show provides an example. A dog show is not a random sampling of the breed: it is typically limited to dogs that are adult, pure-bred, and exemplary. A histogram of dog weights from a show might plausibly be rather complex, like the yellow-orange distribution shown in the illustrations. Suppose we wanted to predict the weight of a dog based on a certain set of characteristics of each dog. One way to do that is to explain the distribution of weights by dividing the dog population into groups based on those characteristics. A successful grouping will split dogs such that (a) each group has a low variance of dog weights (meaning the group is relatively homogeneous) and (b) the mean of each group is distinct (if two groups have the same mean, then it isn't reasonable to conclude that the groups are, in fact, separate in any meaningful way).
In the illustrations to the right, groups are identified as X1, X2, etc. In the first illustration, the dogs are divided according to the product (interaction) of two binary groupings: young vs old, and short-haired vs long-haired (e.g., group 1 is young, short-haired dogs, group 2 is young, long-haired dogs, etc.). Since the distributions of dog weight within each of the groups (shown in blue) has a relatively large variance, and since the means are very similar across groups, grouping dogs by these characteristics does not produce an effective way to explain the variation in dog weights: knowing which group a dog is in doesn't allow us to predict its weight much better than simply knowing the dog is in a dog show. Thus, this grouping fails to explain the variation in the overall distribution (yellow-orange). | [
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634_2 | An attempt to explain the weight distribution by grouping dogs as pet vs working breed and less athletic vs more athletic would probably be somewhat more successful (fair fit). The heaviest show dogs are likely to be big, strong, working breeds, while breeds kept as pets tend to be smaller and thus lighter. As shown by the second illustration, the distributions have variances that are considerably smaller than in the first case, and the means are more distinguishable. However, the significant overlap of distributions, for example, means that we cannot distinguish X1 and X2 reliably. Grouping dogs according to a coin flip might produce distributions that look similar.
An attempt to explain weight by breed is likely to produce a very good fit. All Chihuahuas are light and all St Bernards are heavy. The difference in weights between Setters and Pointers does not justify separate breeds. The analysis of variance provides the formal tools to justify these intuitive judgments. A common use of the method is the analysis of experimental data or the development of models. The method has some advantages over correlation: not all of the data must be numeric and one result of the method is a judgment in the confidence in an explanatory relationship.
Classes of models
There are three classes of models used in the analysis of variance, and these are outlined here.
Fixed-effects models
The fixed-effects model (class I) of analysis of variance applies to situations in which the experimenter applies one or more treatments to the subjects of the experiment to see whether the response variable values change. This allows the experimenter to estimate the ranges of response variable values that the treatment would generate in the population as a whole.
Random-effects models
Random-effects model (class II) is used when the treatments are not fixed. This occurs when the various factor levels are sampled from a larger population. Because the levels themselves are random variables, some assumptions and the method of contrasting the treatments (a multi-variable generalization of simple differences) differ from the fixed-effects model.
Mixed-effects models
A mixed-effects model (class III) contains experimental factors of both fixed and random-effects types, with appropriately different interpretations and analysis for the two types. | [
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634_3 | Example:
Teaching experiments could be performed by a college or university department to find a good introductory textbook, with each text considered a treatment. The fixed-effects model would compare a list of candidate texts. The random-effects model would determine whether important differences exist among a list of randomly selected texts. The mixed-effects model would compare the (fixed) incumbent texts to randomly selected alternatives.
Defining fixed and random effects has proven elusive, with competing definitions arguably leading toward a linguistic quagmire.
Assumptions
The analysis of variance has been studied from several approaches, the most common of which uses a linear model that relates the response to the treatments and blocks. Note that the model is linear in parameters but may be nonlinear across factor levels. Interpretation is easy when data is balanced across factors but much deeper understanding is needed for unbalanced data.
Textbook analysis using a normal distribution
The analysis of variance can be presented in terms of a linear model, which makes the following assumptions about the probability distribution of the responses:
Independence of observations – this is an assumption of the model that simplifies the statistical analysis.
Normality – the distributions of the residuals are normal.
Equality (or "homogeneity") of variances, called homoscedasticity — the variance of data in groups should be the same.
The separate assumptions of the textbook model imply that the errors are independently, identically, and normally distributed for fixed effects models, that is, that the errors () are independent and
Randomization-based analysis
In a randomized controlled experiment, the treatments are randomly assigned to experimental units, following the experimental protocol. This randomization is objective and declared before the experiment is carried out. The objective random-assignment is used to test the significance of the null hypothesis, following the ideas of C. S. Peirce and Ronald Fisher. This design-based analysis was discussed and developed by Francis J. Anscombe at Rothamsted Experimental Station and by Oscar Kempthorne at Iowa State University. Kempthorne and his students make an assumption of unit treatment additivity, which is discussed in the books of Kempthorne and David R. Cox. | [
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634_4 | Unit-treatment additivity
In its simplest form, the assumption of unit-treatment additivity states that the observed response from experimental unit when receiving treatment can be written as the sum of the unit's response and the treatment-effect , that is
The assumption of unit-treatment additivity implies that, for every treatment , the th treatment has exactly the same effect on every experiment unit.
The assumption of unit treatment additivity usually cannot be directly falsified, according to Cox and Kempthorne. However, many consequences of treatment-unit additivity can be falsified. For a randomized experiment, the assumption of unit-treatment additivity implies that the variance is constant for all treatments. Therefore, by contraposition, a necessary condition for unit-treatment additivity is that the variance is constant.
The use of unit treatment additivity and randomization is similar to the design-based inference that is standard in finite-population survey sampling.
Derived linear model
Kempthorne uses the randomization-distribution and the assumption of unit treatment additivity to produce a derived linear model, very similar to the textbook model discussed previously. The test statistics of this derived linear model are closely approximated by the test statistics of an appropriate normal linear model, according to approximation theorems and simulation studies. However, there are differences. For example, the randomization-based analysis results in a small but (strictly) negative correlation between the observations. In the randomization-based analysis, there is no assumption of a normal distribution and certainly no assumption of independence. On the contrary, the observations are dependent!
The randomization-based analysis has the disadvantage that its exposition involves tedious algebra and extensive time. Since the randomization-based analysis is complicated and is closely approximated by the approach using a normal linear model, most teachers emphasize the normal linear model approach. Few statisticians object to model-based analysis of balanced randomized experiments. | [
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] |
634_5 | Statistical models for observational data
However, when applied to data from non-randomized experiments or observational studies, model-based analysis lacks the warrant of randomization. For observational data, the derivation of confidence intervals must use subjective models, as emphasized by Ronald Fisher and his followers. In practice, the estimates of treatment-effects from observational studies generally are often inconsistent. In practice, "statistical models" and observational data are useful for suggesting hypotheses that should be treated very cautiously by the public.
Summary of assumptions
The normal-model based ANOVA analysis assumes the independence, normality and homogeneity of variances of the residuals. The randomization-based analysis assumes only the homogeneity of the variances of the residuals (as a consequence of unit-treatment additivity) and uses the randomization procedure of the experiment. Both these analyses require homoscedasticity, as an assumption for the normal-model analysis and as a consequence of randomization and additivity for the randomization-based analysis.
However, studies of processes that change variances rather than means (called dispersion effects) have been successfully conducted using ANOVA. There are no necessary assumptions for ANOVA in its full generality, but the F-test used for ANOVA hypothesis testing has assumptions and practical
limitations which are of continuing interest.
Problems which do not satisfy the assumptions of ANOVA can often be transformed to satisfy the assumptions.
The property of unit-treatment additivity is not invariant under a "change of scale", so statisticians often use transformations to achieve unit-treatment additivity. If the response variable is expected to follow a parametric family of probability distributions, then the statistician may specify (in the protocol for the experiment or observational study) that the responses be transformed to stabilize the variance. Also, a statistician may specify that logarithmic transforms be applied to the responses, which are believed to follow a multiplicative model.
According to Cauchy's functional equation theorem, the logarithm is the only continuous transformation that transforms real multiplication to addition. | [
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] |
634_6 | Characteristics
ANOVA is used in the analysis of comparative experiments, those in which only the difference in outcomes is of interest. The statistical significance of the experiment is determined by a ratio of two variances. This ratio is independent of several possible alterations to the experimental observations: Adding a constant to all observations does not alter significance. Multiplying all observations by a constant does not alter significance. So ANOVA statistical significance result is independent of constant bias and scaling errors as well as the units used in expressing observations. In the era of mechanical calculation it was common to subtract a constant from all observations (when equivalent to dropping leading digits) to simplify data entry. This is an example of data coding.
Logic
The calculations of ANOVA can be characterized as computing a number of means and variances, dividing two variances and comparing the ratio to a handbook value to determine statistical significance. Calculating a treatment effect is then trivial: "the effect of any treatment is estimated by taking the difference between the mean of the observations which receive the treatment and the general mean".
Partitioning of the sum of squares
ANOVA uses traditional standardized terminology. The definitional equation of sample variance is , where the divisor is called the degrees of freedom (DF), the summation is called
the sum of squares (SS), the result is called the mean square (MS) and the squared terms are deviations from the sample mean. ANOVA estimates 3 sample variances: a total variance based on all the observation deviations from the grand mean, an error variance based on all the observation deviations from their appropriate treatment means, and a treatment variance. The treatment variance is based on the deviations of treatment means from the grand mean, the result being multiplied by the number of observations in each treatment to account for the difference between the variance of observations and the variance of means.
The fundamental technique is a partitioning of the total sum of squares SS into components related to the effects used in the model. For example, the model for a simplified ANOVA with one type of treatment at different levels.
The number of degrees of freedom DF can be partitioned in a similar way: one of these components (that for error) specifies a chi-squared distribution which describes the associated sum of squares, while the same is true for "treatments" if there is no treatment effect. | [
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634_7 | See also Lack-of-fit sum of squares.
The F-test
The F-test is used for comparing the factors of the total deviation. For example, in one-way, or single-factor ANOVA, statistical significance is tested for by comparing the F test statistic
where MS is mean square, is the number of treatments and is the total number of cases
to the F-distribution with , degrees of freedom. Using the F-distribution is a natural candidate because the test statistic is the ratio of two scaled sums of squares each of which follows a scaled chi-squared distribution.
The expected value of F is (where is the treatment sample size) which is 1 for no treatment effect. As values of F increase above 1, the evidence is increasingly inconsistent with the null hypothesis. Two apparent experimental methods of increasing F are increasing the sample size and reducing the error variance by tight experimental controls.
There are two methods of concluding the ANOVA hypothesis test, both of which produce the same result:
The textbook method is to compare the observed value of F with the critical value of F determined from tables. The critical value of F is a function of the degrees of freedom of the numerator and the denominator and the significance level (α). If F ≥ FCritical, the null hypothesis is rejected.
The computer method calculates the probability (p-value) of a value of F greater than or equal to the observed value. The null hypothesis is rejected if this probability is less than or equal to the significance level (α).
The ANOVA F-test is known to be nearly optimal in the sense of minimizing false negative errors for a fixed rate of false positive errors (i.e. maximizing power for a fixed significance level). For example, to test the hypothesis that various medical treatments have exactly the same effect, the F-test's p-values closely approximate the permutation test's p-values: The approximation is particularly close when the design is balanced. Such permutation tests characterize tests with maximum power against all alternative hypotheses, as observed by Rosenbaum. The ANOVA F-test (of the null-hypothesis that all treatments have exactly the same effect) is recommended as a practical test, because of its robustness against many alternative distributions. | [
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634_8 | Extended logic
ANOVA consists of separable parts; partitioning sources of variance and hypothesis testing can be used individually. ANOVA is used to support other statistical tools. Regression is first used to fit more complex models to data, then ANOVA is used to compare models with the objective of selecting simple(r) models that adequately describe the data. "Such models could be fit without any reference to ANOVA, but ANOVA tools could then be used to make some sense of the fitted models, and to test hypotheses about batches of coefficients." "[W]e think of the analysis of variance as a way of understanding and structuring multilevel models—not as an alternative to regression but as a tool for summarizing complex high-dimensional inferences ..."
For a single factor
The simplest experiment suitable for ANOVA analysis is the completely randomized experiment with a single factor. More complex experiments with a single factor involve constraints on randomization and include completely randomized blocks and Latin squares (and variants: Graeco-Latin squares, etc.). The more complex experiments share many of the complexities of multiple factors. A relatively complete discussion of the analysis (models, data summaries, ANOVA table) of the completely randomized experiment is available.
There are some alternatives to conventional one-way analysis of variance, e.g.: Welch's heteroscedastic F test, Welch's heteroscedastic F test with trimmed means and Winsorized variances, Brown-Forsythe test, Alexander-Govern test, James second order test and Kruskal-Wallis test, available in onewaytests R
It is useful to represent each data point in the following form, called a statistical model:
where
i = 1, 2, 3, …, R
j = 1, 2, 3, …, C
μ = overall average (mean)
τj = differential effect (response) associated with the j level of X; this assumes that overall the values of τj add to zero (that is, )
εij = noise or error associated with the particular ij data value | [
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634_9 | That is, we envision an additive model that says every data point can be represented by summing three quantities: the true mean, averaged over all factor levels being investigated, plus an incremental component associated with the particular column (factor level), plus a final component associated with everything else affecting that specific data value.
For multiple factors
ANOVA generalizes to the study of the effects of multiple factors. When the experiment includes observations at all combinations of levels of each factor, it is termed factorial. Factorial experiments are more efficient than a series of single factor experiments and the efficiency grows as the number of factors increases. Consequently, factorial designs are heavily used.
The use of ANOVA to study the effects of multiple factors has a complication. In a 3-way ANOVA with factors x, y and z, the ANOVA model includes terms for the main effects (x, y, z) and terms for interactions (xy, xz, yz, xyz).
All terms require hypothesis tests. The proliferation of interaction terms increases the risk that some hypothesis test will produce a false positive by chance. Fortunately, experience says that high order interactions are rare.
The ability to detect interactions is a major advantage of multiple factor ANOVA. Testing one factor at a time hides interactions, but produces apparently inconsistent experimental results.
Caution is advised when encountering interactions; Test interaction terms first and expand the analysis beyond ANOVA if interactions are found. Texts vary in their recommendations regarding the continuation of the ANOVA procedure after encountering an interaction. Interactions complicate the interpretation of experimental data. Neither the calculations of significance nor the estimated treatment effects can be taken at face value. "A significant interaction will often mask the significance of main effects." Graphical methods are recommended to enhance understanding. Regression is often useful. A lengthy discussion of interactions is available in Cox (1958). Some interactions can be removed (by transformations) while others cannot.
A variety of techniques are used with multiple factor ANOVA to reduce expense. One technique used in factorial designs is to minimize replication (possibly no replication with support of analytical trickery) and to combine groups when effects are found to be statistically (or practically) insignificant. An experiment with many insignificant factors may collapse into one with a few factors supported by many replications. | [
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634_10 | Associated analysis
Some analysis is required in support of the design of the experiment while other analysis is performed after changes in the factors are formally found to produce statistically significant changes in the responses. Because experimentation is iterative, the results of one experiment alter plans for following experiments.
Preparatory analysis
The number of experimental units
In the design of an experiment, the number of experimental units is planned to satisfy the goals of the experiment. Experimentation is often sequential.
Early experiments are often designed to provide mean-unbiased estimates of treatment effects and of experimental error. Later experiments are often designed to test a hypothesis that a treatment effect has an important magnitude; in this case, the number of experimental units is chosen so that the experiment is within budget and has adequate power, among other goals.
Reporting sample size analysis is generally required in psychology. "Provide information on sample size and the process that led to sample size decisions." The analysis, which is written in the experimental protocol before the experiment is conducted, is examined in grant applications and administrative review boards.
Besides the power analysis, there are less formal methods for selecting the number of experimental units. These include graphical methods based on limiting the probability of false negative errors, graphical methods based on an expected variation increase (above the residuals) and methods based on achieving a desired confidence interval.
Power analysis
Power analysis is often applied in the context of ANOVA in order to assess the probability of successfully rejecting the null hypothesis if we assume a certain ANOVA design, effect size in the population, sample size and significance level. Power analysis can assist in study design by determining what sample size would be required in order to have a reasonable chance of rejecting the null hypothesis when the alternative hypothesis is true.
Effect size
Several standardized measures of effect have been proposed for ANOVA to summarize the strength of the association between a predictor(s) and the dependent variable or the overall standardized difference of the complete model. Standardized effect-size estimates facilitate comparison of findings across studies and disciplines. However, while standardized effect sizes are commonly used in much of the professional literature, a non-standardized measure of effect size that has immediately "meaningful" units may be preferable for reporting purposes. | [
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634_11 | Model confirmation
Sometimes tests are conducted to determine whether the assumptions of ANOVA appear to be violated. Residuals are examined or analyzed to confirm homoscedasticity and gross normality. Residuals should have the appearance of (zero mean normal distribution) noise when plotted as a function of anything including time and
modeled data values. Trends hint at interactions among factors or among observations.
Follow-up tests
A statistically significant effect in ANOVA is often followed by additional tests. This can be done in order to assess which groups are different from which other groups or to test various other focused hypotheses. Follow-up tests are often distinguished in terms of whether they are "planned" (a priori) or "post hoc." Planned tests are determined before looking at the data, and post hoc tests are conceived only after looking at the data (though the term "post hoc" is inconsistently used).
The follow-up tests may be "simple" pairwise comparisons of individual group means or may be "compound" comparisons (e.g., comparing the mean pooling across groups A, B and C to the mean of group D). Comparisons can also look at tests of trend, such as linear and quadratic relationships, when the independent variable involves ordered levels. Often the follow-up tests incorporate a method of adjusting for the multiple comparisons problem.
Study designs
There are several types of ANOVA. Many statisticians base ANOVA on the design of the experiment, especially on the protocol that specifies the random assignment of treatments to subjects; the protocol's description of the assignment mechanism should include a specification of the structure of the treatments and of any blocking. It is also common to apply ANOVA to observational data using an appropriate statistical model. | [
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634_12 | Some popular designs use the following types of ANOVA:
One-way ANOVA is used to test for differences among two or more independent groups (means), e.g. different levels of urea application in a crop, or different levels of antibiotic action on several different bacterial species, or different levels of effect of some medicine on groups of patients. However, should these groups not be independent, and there is an order in the groups (such as mild, moderate and severe disease), or in the dose of a drug (such as 5 mg/mL, 10 mg/mL, 20 mg/mL) given to the same group of patients, then a linear trend estimation should be used. Typically, however, the one-way ANOVA is used to test for differences among at least three groups, since the two-group case can be covered by a t-test. When there are only two means to compare, the t-test and the ANOVA F-test are equivalent; the relation between ANOVA and t is given by .
Factorial ANOVA is used when there is more than one factor.
Repeated measures ANOVA is used when the same subjects are used for each factor (e.g., in a longitudinal study).
Multivariate analysis of variance (MANOVA) is used when there is more than one response variable.
Cautions
Balanced experiments (those with an equal sample size for each treatment) are relatively easy to interpret; unbalanced experiments offer more complexity. For single-factor (one-way) ANOVA, the adjustment for unbalanced data is easy, but the unbalanced analysis lacks both robustness and power. For more complex designs the lack of balance leads to further complications. "The orthogonality property of main effects and interactions present in balanced data does not carry over to the unbalanced case. This means that the usual analysis of variance techniques do not apply. Consequently, the analysis of unbalanced factorials is much more difficult than that for balanced designs." In the general case, "The analysis of variance can also be applied to unbalanced data, but then the sums of squares, mean squares, and F-ratios will depend on the order in which the sources of variation are considered."
ANOVA is (in part) a test of statistical significance. The American Psychological Association (and many other organisations) holds the view that simply reporting statistical significance is insufficient and that reporting confidence bounds is preferred. | [
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] |
634_13 | Generalizations
ANOVA is considered to be a special case of linear regression which in turn is a special case of the general linear model. All consider the observations to be the sum of a model (fit) and a residual (error) to be minimized.
The Kruskal–Wallis test and the Friedman test are nonparametric tests, which do not rely on an assumption of normality.
Connection to linear regression
Below we make clear the connection between multi-way ANOVA and linear regression.
Linearly re-order the data so that -th observation is associated with a response and factors where denotes the different factors and is the total number of factors. In one-way ANOVA and in two-way ANOVA . Furthermore, we assume the -th factor has levels, namely . Now, we can one-hot encode the factors into the dimensional vector .
The one-hot encoding function is defined such that the -th entry of is
The vector is the concatenation of all of the above vectors for all . Thus, . In order to obtain a fully general -way interaction ANOVA we must also concatenate every additional interaction term in the vector and then add an intercept term. Let that vector be .
With this notation in place, we now have the exact connection with linear regression. We simply regress response against the vector . However, there is a concern about identifiability. In order to overcome such issues we assume that the sum of the parameters within each set of interactions is equal to zero. From here, one can use F-statistics or other methods to determine the relevance of the individual factors.
Example
We can consider the 2-way interaction example where we assume that the first factor has 2 levels and the second factor has 3 levels.
Define if and if , i.e. is the one-hot encoding of the first factor and is the one-hot encoding of the second factor.
With that,
where the last term is an intercept term. For a more concrete example suppose that
Then,
See also | [
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] |
634_14 | Then,
See also
ANOVA on ranks
ANOVA-simultaneous component analysis
Analysis of covariance (ANCOVA)
Analysis of molecular variance (AMOVA)
Analysis of rhythmic variance (ANORVA)
Explained variation
Linear trend estimation
Mixed-design analysis of variance
Multivariate analysis of covariance (MANCOVA)
Permutational analysis of variance
Variance decomposition
Expected mean squares
Footnotes
Notes
References
Pre-publication chapters are available on-line.
Cohen, Jacob (1988). Statistical power analysis for the behavior sciences (2nd ed.). Routledge
Cox, David R. (1958). Planning of experiments. Reprinted as
Freedman, David A.(2005). Statistical Models: Theory and Practice, Cambridge University Press.
Lehmann, E.L. (1959) Testing Statistical Hypotheses. John Wiley & Sons.
Moore, David S. & McCabe, George P. (2003). Introduction to the Practice of Statistics (4e). W H Freeman & Co.
Rosenbaum, Paul R. (2002). Observational Studies (2nd ed.). New York: Springer-Verlag.
Further reading
Cox, David R. & Reid, Nancy M. (2000). The theory of design of experiments. (Chapman & Hall/CRC).
Freedman, David A.; Pisani, Robert; Purves, Roger (2007) Statistics, 4th edition. W.W. Norton & Company
Tabachnick, Barbara G. & Fidell, Linda S. (2007). Using Multivariate Statistics (5th ed.). Boston: Pearson International Edition.
External links
SOCR ANOVA Activity
Examples of all ANOVA and ANCOVA models with up to three treatment factors, including randomized block, split plot, repeated measures, and Latin squares, and their analysis in R (University of Southampton)
NIST/SEMATECH e-Handbook of Statistical Methods, section 7.4.3: "Are the means equal?"
Analysis of variance: Introduction
Design of experiments
Statistical tests
Parametric statistics | [
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639_0 | In organic chemistry, an alkane, or paraffin (a historical trivial name that also has other meanings), is an acyclic saturated hydrocarbon. In other words, an alkane consists of hydrogen and carbon atoms arranged in a tree structure in which all the carbon–carbon bonds are single. Alkanes have the general chemical formula . The alkanes range in complexity from the simplest case of methane (), where n = 1 (sometimes called the parent molecule), to arbitrarily large and complex molecules, like pentacontane () or 6-ethyl-2-methyl-5-(1-methylethyl) octane, an isomer of tetradecane ().
The International Union of Pure and Applied Chemistry (IUPAC) defines alkanes as "acyclic branched or unbranched hydrocarbons having the general formula , and therefore consisting entirely of hydrogen atoms and saturated carbon atoms". However, some sources use the term to denote any saturated hydrocarbon, including those that are either monocyclic (i.e. the cycloalkanes) or polycyclic, despite their having a distinct general formula (i.e. cycloalkanes are ).
In an alkane, each carbon atom is sp3-hybridized with 4 sigma bonds (either C–C or C–H), and each hydrogen atom is joined to one of the carbon atoms (in a C–H bond). The longest series of linked carbon atoms in a molecule is known as its carbon skeleton or carbon backbone. The number of carbon atoms may be considered as the size of the alkane.
One group of the higher alkanes are waxes, solids at standard ambient temperature and pressure (SATP), for which the number of carbon atoms in the carbon backbone is greater than about 17.
With their repeated – units, the alkanes constitute a homologous series of organic compounds in which the members differ in molecular mass by multiples of 14.03 u (the total mass of each such methylene-bridge unit, which comprises a single carbon atom of mass 12.01 u and two hydrogen atoms of mass ~1.01 u each). | [
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639_1 | Methane is produced by methanogenic bacteria and some long-chain alkanes function as pheromones in certain animal species or as protective waxes in plants and fungi. Nevertheless, most alkanes do not have much biological activity. They can be viewed as molecular trees upon which can be hung the more active/reactive functional groups of biological molecules.
The alkanes have two main commercial sources: petroleum (crude oil) and natural gas.
An alkyl group is an alkane-based molecular fragment that bears one open valence for bonding. They are generally abbreviated with the symbol for any organyl group, R, although Alk is sometimes used to specifically symbolize an alkyl group (as opposed to an alkenyl group or aryl group).
Structure and classification
Ordinarily the C-C single bond distance is .
Saturated hydrocarbons can be linear, branched, or cyclic. The third group is sometimes called cycloalkanes. Very complicated structures are possible by combining linear, branch, cyclic alkanes.
Isomerism | [
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639_2 | Isomerism
Alkanes with more than three carbon atoms can be arranged in various ways, forming structural isomers. The simplest isomer of an alkane is the one in which the carbon atoms are arranged in a single chain with no branches. This isomer is sometimes called the n-isomer (n for "normal", although it is not necessarily the most common). However, the chain of carbon atoms may also be branched at one or more points. The number of possible isomers increases rapidly with the number of carbon atoms. For example, for acyclic alkanes:
C1: methane only
C2: ethane only
C3: propane only
C4: 2 isomers: butane and isobutane
C5: 3 isomers: pentane, isopentane, and neopentane
C6: 5 isomers: hexane, 2-methylpentane, 3-methylpentane, 2,2-dimethylbutane, and 2,3-dimethylbutane
C7: 9 isomers: heptane, methylhexane (2 isomers), dimethylpentane (4 isomers), 3-ethylpentane, 2,2,3-trimethylbutane
C8: 18 isomers: octane, 2-methylheptane, 3-methylheptane, 2,3-dimethylhexane, 3,4-dimethylhexane, 2,3,4-trimethylpentane, 3,3-dimethylhexane, 2,2-trimethylpentane, 2,4-dimethylhexane, 2,2,4-trimethylpentane, 2,3,3-Trimethylpentane, 3,3,4-trimethyl-pentane, 3,4,4-trimethylpentane, 2,4,4-trimethylpentane, (5 isomers)
C9: 35 isomers
C10: 75 isomers
C12: 355 isomers
C32: 27,711,253,769 isomers
C60: 22,158,734,535,770,411,074,184 isomers, many of which are not stable. | [
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639_3 | Branched alkanes can be chiral. For example, 3-methylhexane and its higher homologues are chiral due to their stereogenic center at carbon atom number 3. The above list only includes differences of connectivity, not stereochemistry. In addition to the alkane isomers, the chain of carbon atoms may form one or more rings. Such compounds are called cycloalkanes, and are also excluded from the above list because changing the number of rings changes the molecular formula. For example, cyclobutane and methylcyclopropane are isomers of each other (C4H8), but are not isomers of butane (C4H10).
Nomenclature
The IUPAC nomenclature (systematic way of naming compounds) for alkanes is based on identifying hydrocarbon chains. Unbranched, saturated hydrocarbon chains are named systematically with a Greek numerical prefix denoting the number of carbons and the suffix "-ane".
In 1866, August Wilhelm von Hofmann suggested systematizing nomenclature by using the whole sequence of vowels a, e, i, o and u to create suffixes -ane, -ene, -ine (or -yne), -one, -une, for the hydrocarbons CnH2n+2, CnH2n, CnH2n−2, CnH2n−4, CnH2n−6. In modern nomenclature, the first three specifically name hydrocarbons with single, double and triple bonds; while "-one" now represents a ketone.
Linear alkanes
Straight-chain alkanes are sometimes indicated by the prefix "n-" or "n-"(for "normal") where a non-linear isomer exists. Although this is not strictly necessary and is not part of the IUPAC naming system, the usage is still common in cases where one wishes to emphasize or distinguish between the straight-chain and branched-chain isomers, e.g., "n-butane" rather than simply "butane" to differentiate it from isobutane. Alternative names for this group used in the petroleum industry are linear paraffins or n-paraffins. | [
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] |
639_4 | The first six members of the series (in terms of number of carbon atoms) are named as follows:
methane CH4 – one carbon and 4 hydrogen
ethane C2H6 – two carbon and 6 hydrogen
propane C3H8 – three carbon and 8 hydrogen
butane C4H10 – four carbon and 10 hydrogen
pentane C5H12 – five carbon and 12 hydrogen
hexane C6H14 – six carbon and 14 hydrogen
The first four names were derived from methanol, ether, propionic acid and butyric acid. Alkanes with five or more carbon atoms are named by adding the suffix -ane to the appropriate numerical multiplier prefix with elision of any terminal vowel (-a or -o) from the basic numerical term. Hence, pentane, C5H12; hexane, C6H14; heptane, C7H16; octane, C8H18; etc. The numeral prefix is generally Greek, however alkanes with a carbon atom count ending in nine, for example nonane, use the Latin prefix non-. For a more complete list, see list of straight-chain alkanes.
Branched alkanes
Simple branched alkanes often have a common name using a prefix to distinguish them from linear alkanes, for example n-pentane, isopentane, and neopentane.
IUPAC naming conventions can be used to produce a systematic name.
The key steps in the naming of more complicated branched alkanes are as follows:
Identify the longest continuous chain of carbon atoms
Name this longest root chain using standard naming rules
Name each side chain by changing the suffix of the name of the alkane from "-ane" to "-yl"
Number the longest continuous chain in order to give the lowest possible numbers for the side-chains
Number and name the side chains before the name of the root chain
If there are multiple side chains of the same type, use prefixes such as "di-" and "tri-" to indicate it as such, and number each one.
Add side chain names in alphabetical (disregarding "di-" etc. prefixes) order in front of the name of the root chain
Saturated cyclic hydrocarbons | [
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] |
639_5 | Saturated cyclic hydrocarbons
Though technically distinct from the alkanes, this class of hydrocarbons is referred to by some as the "cyclic alkanes." As their description implies, they contain one or more rings.
Simple cycloalkanes have a prefix "cyclo-" to distinguish them from alkanes. Cycloalkanes are named as per their acyclic counterparts with respect to the number of carbon atoms in their backbones, e.g., cyclopentane (C5H10) is a cycloalkane with 5 carbon atoms just like pentane (C5H12), but they are joined up in a five-membered ring. In a similar manner, propane and cyclopropane, butane and cyclobutane, etc.
Substituted cycloalkanes are named similarly to substituted alkanes – the cycloalkane ring is stated, and the substituents are according to their position on the ring, with the numbering decided by the Cahn–Ingold–Prelog priority rules.
Trivial/common names
The trivial (non-systematic) name for alkanes is 'paraffins'. Together, alkanes are known as the 'paraffin series'. Trivial names for compounds are usually historical artifacts. They were coined before the development of systematic names, and have been retained due to familiar usage in industry. Cycloalkanes are also called naphthenes.
Branched-chain alkanes are called isoparaffins. "Paraffin" is a general term and often does not distinguish between pure compounds and mixtures of isomers, i.e., compounds of the same chemical formula, e.g., pentane and isopentane.
In IUPAC
The following trivial names are retained in the IUPAC system:
isobutane for 2-methylpropane
isopentane for 2-methylbutane
neopentane for 2,2-dimethylpropane.
Non-IUPAC
Some non-IUPAC trivial names are occasionally used:
cetane, for hexadecane
cerane, for hexacosane | [
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639_6 | Physical properties
All alkanes are colorless. Alkanes with the lowest molecular weights are gasses, those of intermediate molecular weight are liquids, and the heaviest are waxy solids.
Table of alkanes
Boiling point
Alkanes experience intermolecular van der Waals forces. Stronger intermolecular van der Waals forces give rise to greater boiling points of alkanes.
There are two determinants for the strength of the van der Waals forces:
the number of electrons surrounding the molecule, which increases with the alkane's molecular weight
the surface area of the molecule
Under standard conditions, from CH4 to C4H10 alkanes are gaseous; from C5H12 to C17H36 they are liquids; and after C18H38 they are solids. As the boiling point of alkanes is primarily determined by weight, it should not be a surprise that the boiling point has almost a linear relationship with the size (molecular weight) of the molecule. As a rule of thumb, the boiling point rises 20–30 °C for each carbon added to the chain; this rule applies to other homologous series.
A straight-chain alkane will have a boiling point higher than a branched-chain alkane due to the greater surface area in contact, thus the greater van der Waals forces, between adjacent molecules. For example, compare isobutane (2-methylpropane) and n-butane (butane), which boil at −12 and 0 °C, and 2,2-dimethylbutane and 2,3-dimethylbutane which boil at 50 and 58 °C, respectively.
On the other hand, cycloalkanes tend to have higher boiling points than their linear counterparts due to the locked conformations of the molecules, which give a plane of intermolecular contact. | [
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639_7 | Melting points
The melting points of the alkanes follow a similar trend to boiling points for the same reason as outlined above. That is, (all other things being equal) the larger the molecule the higher the melting point. There is one significant difference between boiling points and melting points. Solids have more rigid and fixed structure than liquids. This rigid structure requires energy to break down. Thus the better put together solid structures will require more energy to break apart. For alkanes, this can be seen from the graph above (i.e., the blue line). The odd-numbered alkanes have a lower trend in melting points than even numbered alkanes. This is because even numbered alkanes pack well in the solid phase, forming a well-organized structure, which requires more energy to break apart. The odd-numbered alkanes pack less well and so the "looser" organized solid packing structure requires less energy to break apart. For a visualization of the crystal structures see.
The melting points of branched-chain alkanes can be either higher or lower than those of the corresponding straight-chain alkanes, again depending on the ability of the alkane in question to pack well in the solid phase.
Conductivity and solubility
Alkanes do not conduct electricity in any way, nor are they substantially polarized by an electric field. For this reason, they do not form hydrogen bonds and are insoluble in polar solvents such as water. Since the hydrogen bonds between individual water molecules are aligned away from an alkane molecule, the coexistence of an alkane and water leads to an increase in molecular order (a reduction in entropy). As there is no significant bonding between water molecules and alkane molecules, the second law of thermodynamics suggests that this reduction in entropy should be minimized by minimizing the contact between alkane and water: Alkanes are said to be hydrophobic as they are insoluble in water.
Their solubility in nonpolar solvents is relatively high, a property that is called lipophilicity. Alkanes are, for example, miscible in all proportions among themselves. | [
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639_8 | The density of the alkanes usually increases with the number of carbon atoms but remains less than that of water. Hence, alkanes form the upper layer in an alkane–water mixture.
Molecular geometry
The molecular structure of the alkanes directly affects their physical and chemical characteristics. It is derived from the electron configuration of carbon, which has four valence electrons. The carbon atoms in alkanes are described as sp3 hybrids, that is to say that, to a good approximation, the valence electrons are in orbitals directed towards the corners of a tetrahedron which are derived from the combination of the 2s orbital and the three 2p orbitals. Geometrically, the angle between the bonds are cos−1(−) ≈ 109.47°. This is exact for the case of methane, while larger alkanes containing a combination of C–H and C–C bonds generally have bonds that are within several degrees of this idealized value.
Bond lengths and bond angles
An alkane has only C–H and C–C single bonds. The former result from the overlap of an sp3 orbital of carbon with the 1s orbital of a hydrogen; the latter by the overlap of two sp3 orbitals on adjacent carbon atoms. The bond lengths amount to 1.09 × 10−10 m for a C–H bond and 1.54 × 10−10 m for a C–C bond.
The spatial arrangement of the bonds is similar to that of the four sp3 orbitals—they are tetrahedrally arranged, with an angle of 109.47° between them. Structural formulae that represent the bonds as being at right angles to one another, while both common and useful, do not accurately depict the geometry.
Conformation
The structural formula and the bond angles are not usually sufficient to completely describe the geometry of a molecule. There is a further degree of freedom for each carbon–carbon bond: the torsion angle between the atoms or groups bound to the atoms at each end of the bond. The spatial arrangement described by the torsion angles of the molecule is known as its conformation. | [
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639_9 | Ethane forms the simplest case for studying the conformation of alkanes, as there is only one C–C bond. If one looks down the axis of the C–C bond, one will see the so-called Newman projection. The hydrogen atoms on both the front and rear carbon atoms have an angle of 120° between them, resulting from the projection of the base of the tetrahedron onto a flat plane. However, the torsion angle between a given hydrogen atom attached to the front carbon and a given hydrogen atom attached to the rear carbon can vary freely between 0° and 360°. This is a consequence of the free rotation about a carbon–carbon single bond. Despite this apparent freedom, only two limiting conformations are important: eclipsed conformation and staggered conformation.
The two conformations differ in energy: the staggered conformation is 12.6 kJ/mol (3.0 kcal/mol) lower in energy (more stable) than the eclipsed conformation (the least stable).
This difference in energy between the two conformations, known as the torsion energy, is low compared to the thermal energy of an ethane molecule at ambient temperature. There is constant rotation about the C–C bond. The time taken for an ethane molecule to pass from one staggered conformation to the next, equivalent to the rotation of one CH3 group by 120° relative to the other, is of the order of 10−11 seconds.
The case of higher alkanes is more complex but based on similar principles, with the antiperiplanar conformation always being the most favored around each carbon–carbon bond. For this reason, alkanes are usually shown in a zigzag arrangement in diagrams or in models. The actual structure will always differ somewhat from these idealized forms, as the differences in energy between the conformations are small compared to the thermal energy of the molecules: Alkane molecules have no fixed structural form, whatever the models may suggest.
Spectroscopic properties
Virtually all organic compounds contain carbon–carbon, and carbon–hydrogen bonds, and so show some of the features of alkanes in their spectra. Alkanes are notable for having no other groups, and therefore for the absence of other characteristic spectroscopic features of a functional group like –OH, –CHO, –COOH etc. | [
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] |
639_10 | Infrared spectroscopy
The carbon–hydrogen stretching mode gives a strong absorption between 2850 and 2960 cm−1, while the carbon–carbon stretching mode absorbs between 800 and 1300 cm−1. The carbon–hydrogen bending modes depend on the nature of the group: methyl groups show bands at 1450 cm−1 and 1375 cm−1, while methylene groups show bands at 1465 cm−1 and 1450 cm−1. Carbon chains with more than four carbon atoms show a weak absorption at around 725 cm−1.
NMR spectroscopy
The proton resonances of alkanes are usually found at δH = 0.5–1.5. The carbon-13 resonances depend on the number of hydrogen atoms attached to the carbon: δC = 8–30 (primary, methyl, –CH3), 15–55 (secondary, methylene, –CH2–), 20–60 (tertiary, methyne, C–H) and quaternary. The carbon-13 resonance of quaternary carbon atoms is characteristically weak, due to the lack of nuclear Overhauser effect and the long relaxation time, and can be missed in weak samples, or samples that have not been run for a sufficiently long time.
Mass spectrometry
Alkanes have a high ionization energy, and the molecular ion is usually weak. The fragmentation pattern can be difficult to interpret, but, in the case of branched chain alkanes, the carbon chain is preferentially cleaved at tertiary or quaternary carbons due to the relative stability of the resulting free radicals. The fragment resulting from the loss of a single methyl group (M − 15) is often absent, and other fragments are often spaced by intervals of fourteen mass units, corresponding to sequential loss of CH2 groups.
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